<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD Journal Publishing DTD v3.0 20080202//EN" "https://jats.nlm.nih.gov/nlm-dtd/publishing/3.0/journalpublishing3.dtd">
<article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" article-type="research-article" specific-use="SMUR" dtd-version="3.0" xml:lang="en">
<front>
<journal-meta>
<journal-id journal-id-type="publisher">ESDD</journal-id>
<journal-title-group>
<journal-title>Earth System Dynamics Discussions</journal-title>
<abbrev-journal-title abbrev-type="publisher">ESDD</abbrev-journal-title>
<abbrev-journal-title abbrev-type="nlm-ta">Earth Syst. Dynam. Discuss.</abbrev-journal-title>
</journal-title-group>
<issn pub-type="epub">2190-4995</issn>
<publisher><publisher-name></publisher-name>
<publisher-loc>Göttingen, Germany</publisher-loc>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="doi">10.5194/esdd-2-211-2011</article-id>
<title-group>
<article-title>Geologic constraints on earth system sensitivity to CO&lt;sub&gt;2&lt;/sub&gt; during the Cretaceous and early Paleogene</article-title>
</title-group>
<contrib-group><contrib contrib-type="author" xlink:type="simple"><name name-style="western"><surname>Royer</surname>
<given-names>D. L.</given-names>
</name>
<xref ref-type="aff" rid="aff1">
<sup>1</sup>
</xref>
</contrib>
<contrib contrib-type="author" xlink:type="simple"><name name-style="western"><surname>Pagani</surname>
<given-names>M.</given-names>
</name>
<xref ref-type="aff" rid="aff2">
<sup>2</sup>
</xref>
</contrib>
<contrib contrib-type="author" xlink:type="simple"><name name-style="western"><surname>Beerling</surname>
<given-names>D. J.</given-names>
</name>
<xref ref-type="aff" rid="aff3">
<sup>3</sup>
</xref>
</contrib>
</contrib-group><aff id="aff1">
<label>1</label>
<addr-line>Department of Earth and Environmental Sciences and College of the Environment, Wesleyan University, Middletown, Connecticut, 06459, USA</addr-line>
</aff>
<aff id="aff2">
<label>2</label>
<addr-line>Department of Geology and Geophysics, Yale University, New Haven, Connecticut, 06520, USA</addr-line>
</aff>
<aff id="aff3">
<label>3</label>
<addr-line>Department of Animal and Plant Sciences, University of Sheffield, Sheffield, S10 2TN, UK</addr-line>
</aff>
<pub-date pub-type="epub">
<day>03</day>
<month>03</month>
<year>2011</year>
</pub-date>
<volume>2</volume>
<issue>1</issue>
<fpage>211</fpage>
<lpage>240</lpage>
<permissions>
<copyright-statement>Copyright: &#x000a9; 2011 D. L. Royer et al.</copyright-statement>
<copyright-year>2011</copyright-year>
<license license-type="open-access">
<license-p>This work is licensed under the Creative Commons Attribution 3.0 Unported License. To view a copy of this licence, visit <ext-link ext-link-type="uri"  xlink:href="https://creativecommons.org/licenses/by/3.0/">https://creativecommons.org/licenses/by/3.0/</ext-link></license-p>
</license>
</permissions>
<self-uri xlink:href="https://esd.copernicus.org/preprints/2/211/2011/esdd-2-211-2011.html">This article is available from https://esd.copernicus.org/preprints/2/211/2011/esdd-2-211-2011.html</self-uri>
<self-uri xlink:href="https://esd.copernicus.org/preprints/2/211/2011/esdd-2-211-2011.pdf">The full text article is available as a PDF file from https://esd.copernicus.org/preprints/2/211/2011/esdd-2-211-2011.pdf</self-uri>
<abstract>
<p>Earth system sensitivity (ESS) is the long-term (&gt;10&lt;sup&gt;3&lt;/sup&gt; yr) equilibrium temperature
response to doubled CO&lt;sub&gt;2&lt;/sub&gt;. ESS has climate policy implications because
global temperatures are not expected to decline appreciably for at least
10&lt;sup&gt;3&lt;/sup&gt; yr, even if anthropogenic greenhouse-gas emissions drop to zero. We
report quantitative ESS estimates of 3 °C or higher for much of the
Cretaceous and early Paleogene based on paleo-reconstructions of CO&lt;sub&gt;2&lt;/sub&gt;
and temperature. These estimates are generally higher than climate
sensitivities simulated from global climate models for the same ancient
periods (~3 °C). We conclude that climate models do not capture
the full suite of positive climate feedbacks during greenhouse worlds. These
absent feedbacks are probably related to clouds, trace greenhouse gases,
seasonal snow cover, and/or vegetation, especially in polar regions. Continued
warming in the coming decades as anthropogenic greenhouse gases accumulate
in the atmosphere ensures that characterizing and quantifying these positive
climate feedbacks will become a scientific challenge of increasing priority.</p>
</abstract>
<counts><page-count count="30"/></counts>
</article-meta>
</front>
<body/>
<back>
<ref-list>
<title>References</title>
<ref id="ref1">
<label>1</label><mixed-citation publication-type="other" xlink:type="simple">Archer, D.: Fate of fossil fuel CO&lt;sub&gt;2&lt;/sub&gt; in geologic time, J. Geophys. Res., 110, C09S05, &lt;a href=&quot;http://dx.doi.org/10.1029/2004JC002625&quot;&gt;https://doi.org/10.1029/2004JC002625&lt;/a&gt;, 2005.</mixed-citation>
</ref>
<ref id="ref2">
<label>2</label><mixed-citation publication-type="other" xlink:type="simple">Archer, D. and Brovkin, V.: The millennial atmospheric lifetime of anthropogenic CO&lt;sub&gt;2&lt;/sub&gt;, Climatic Change, 90, 283–297, 2008.</mixed-citation>
</ref>
<ref id="ref3">
<label>3</label><mixed-citation publication-type="other" xlink:type="simple">Armour, K. C. and Roe, G. H.: Climate commitment in an uncertain world, Geophys. Res. Lett., 38, L01707, &lt;a href=&quot;http://dx.doi.org/10.1029/2010GL045850&quot;&gt;https://doi.org/10.1029/2010GL045850&lt;/a&gt;, 2011.</mixed-citation>
</ref>
<ref id="ref4">
<label>4</label><mixed-citation publication-type="other" xlink:type="simple">Arneth, A., Harrison, S. P., Zaehle, S., Tsigaridis, K., Menon, S., Bartlein, P. J., and Feichter, J.: Terrestrial biogeochemical feedbacks in the climate system, Nat. Geosci., 3, 525–532, 2010.</mixed-citation>
</ref>
<ref id="ref5">
<label>5</label><mixed-citation publication-type="other" xlink:type="simple">Barron, E. J., Fawcett, P. J., Pollard, D., and Thompson, S.: Model simulations of Cretaceous climates: the role of geography and carbon dioxide, Philos. T. Roy. Soc.&amp;nbsp;B, 341, 307–316, 1993.</mixed-citation>
</ref>
<ref id="ref6">
<label>6</label><mixed-citation publication-type="other" xlink:type="simple">Beerling, D. J., Lomax, B. H., Royer, D. L., Upchurch, G. R., and Kump, L. R.: An atmospheric &lt;i&gt;p&lt;/i&gt;CO&lt;sub&gt;2&lt;/sub&gt; reconstruction across the Cretaceous-Tertiary boundary from leaf megafossils, P. Natl. Acad. Sci. USA, 99, 7836–7840, 2002.</mixed-citation>
</ref>
<ref id="ref7">
<label>7</label><mixed-citation publication-type="other" xlink:type="simple">Beerling, D. J., Berner, R. A., Mackenzie, F. T., Harfoot, M. B., and Pyle, J. A.: Methane and the CH&lt;sub&gt;4&lt;/sub&gt;-related greenhouse effect over the past 400 million years, Am. J. Sci., 309, 97–113, 2009a.</mixed-citation>
</ref>
<ref id="ref8">
<label>8</label><mixed-citation publication-type="other" xlink:type="simple">Beerling, D. J., Fox, A., and Anderson, C. W.: Quantitative uncertainty analyses of ancient atmospheric CO&lt;sub&gt;2&lt;/sub&gt; estimates from fossil leaves, Am. J. Sci., 309, 775–787, 2009b.</mixed-citation>
</ref>
<ref id="ref9">
<label>9</label><mixed-citation publication-type="other" xlink:type="simple">Beerling, D. J., Fox, A., Stevenson, D. S., and Valdes, P. J.: Enhanced chemistry-climate feedbacks in past greenhouse worlds, P. Natl. Acad. Sci. USA, in review, 2011.</mixed-citation>
</ref>
<ref id="ref10">
<label>10</label><mixed-citation publication-type="other" xlink:type="simple">Bice, K. L. and Norris, R. D.: Possible atmospheric CO&lt;sub&gt;2&lt;/sub&gt; extremes of the Middle Cretaceous (late Albian-Turonian), Paleoceanography, 17(4), 1070, &lt;a href=&quot;http://dx.doi.org/10.1029/2002PA000778&quot;&gt;https://doi.org/10.1029/2002PA000778&lt;/a&gt;, 2002.</mixed-citation>
</ref>
<ref id="ref11">
<label>11</label><mixed-citation publication-type="other" xlink:type="simple">Bice, K. L., Scotese, C. R., Seidov, D., and Barron, E. J.: Quantifying the role of geographic change in Cenozoic ocean heat transport using uncoupled atmosphere and ocean models, Palaeogeogr. Palaeocl., 161, 295–310, 2000.</mixed-citation>
</ref>
<ref id="ref12">
<label>12</label><mixed-citation publication-type="other" xlink:type="simple">Bice, K. L., Birgel, D., Metyers, P. A., Dahl, K. A., Hinrichs, K.-U., and Norris, R. D.: A multiple proxy and model study of Cretaceous upper ocean temperatures and atmospheric CO&lt;sub&gt;2&lt;/sub&gt; concentration, Paleoceanography, 21, PA2002, &lt;a href=&quot;http://dx.doi.org/10.1029/2005PA001203&quot;&gt;https://doi.org/10.1029/2005PA001203&lt;/a&gt;, 2006.</mixed-citation>
</ref>
<ref id="ref13">
<label>13</label><mixed-citation publication-type="other" xlink:type="simple">Bijl, P. K., Schouten, S., Sluijs, A., Reichart, G.-J., Zachos, J. C., and Brinkhuis, H.: Early Palaeogene temperature evolution of the southwest Pacific Ocean, Nature, 461, 776–779, 2009.</mixed-citation>
</ref>
<ref id="ref14">
<label>14</label><mixed-citation publication-type="other" xlink:type="simple">Bijl, P. K., Houben, A. J. P., Schouten, S., Bohaty, S. M., Sluijs, A., Reichart, G.-J., Sinninghe Damsté, J. S., and Brinkhuis, H.: Transient Middle Eocene atmospheric CO&lt;sub&gt;2&lt;/sub&gt; and temperature variations, Science, 330, 819–821, 2010.</mixed-citation>
</ref>
<ref id="ref15">
<label>15</label><mixed-citation publication-type="other" xlink:type="simple">Bornemann, A., Norris, R. D., Friedrich, O., Beckmann, B., Schouten, S., Sinninghe Damsté, J. S., Vogel, J., Hofmann, P., and Wagner, T.: Isotopic evidence for glaciation during the Cretaceous supergreenhouse, Science, 319, 189–192, 2008.</mixed-citation>
</ref>
<ref id="ref16">
<label>16</label><mixed-citation publication-type="other" xlink:type="simple">Borzenkova, I. I.: Determination of global climate sensitivity to the gas composition of the atmosphere from paleoclimatic data, Izv. Atmos. Ocean. Phys., 39, 197–202, 2003.</mixed-citation>
</ref>
<ref id="ref17">
<label>17</label><mixed-citation publication-type="other" xlink:type="simple">Breecker, D. O., Sharp, Z. D., and McFadden, L. D.: Seasonal bias in the formation and stable isotopic composition of pedogenic carbonate in modern soils from central New Mexico, USA, Geol. Soc. Am. Bull., 121, 630–640, 2009.</mixed-citation>
</ref>
<ref id="ref18">
<label>18</label><mixed-citation publication-type="other" xlink:type="simple">Breecker, D. O., Sharp, Z. D., and McFadden, L. D.: Atmospheric CO&lt;sub&gt;2&lt;/sub&gt; concentrations during ancient greenhouse climates were similar to those predicted for A.D.&amp;nbsp;2100, P. Natl. Acad. Sci. USA, 107, 576–580, 2010.</mixed-citation>
</ref>
<ref id="ref19">
<label>19</label><mixed-citation publication-type="other" xlink:type="simple">Brierley, C. M. and Fedorov, A. V.: Relative importance of meridional and zonal sea surface temperature gradients for the onset of the ice ages and Pliocene-Pleistocene climate evolution, Paleoceanography, 25, PA2214, &lt;a href=&quot;http://dx.doi.org/10.1029/2009PA001809&quot;&gt;https://doi.org/10.1029/2009PA001809&lt;/a&gt;, 2010.</mixed-citation>
</ref>
<ref id="ref20">
<label>20</label><mixed-citation publication-type="other" xlink:type="simple">Budyko, M. I., Ronov, A. B., and Yanshin, A. L.: History of the Earth&apos;s Atmosphere, Springer-Verlag, Berlin, 1987.</mixed-citation>
</ref>
<ref id="ref21">
<label>21</label><mixed-citation publication-type="other" xlink:type="simple">Carslaw, K. S., Boucher, O., Spracklen, D. V., Mann, G. W., Rae, J. G. L., Woodward, S., and Kulmala, M.: A review of natural aerosol interactions and feedbacks within the Earth system, Atmos. Chem. Phys., 10, 1701–1737, &lt;a href=&quot;http://dx.doi.org/10.5194/acp-10-1701-2010&quot;&gt;https://doi.org/10.5194/acp-10-1701-2010&lt;/a&gt;, 2010.</mixed-citation>
</ref>
<ref id="ref22">
<label>22</label><mixed-citation publication-type="other" xlink:type="simple">Charney, J.: Carbon dioxide and climate: a scientific assessment, National Academy of Sciences Press, Washington, DC, 1979.</mixed-citation>
</ref>
<ref id="ref23">
<label>23</label><mixed-citation publication-type="other" xlink:type="simple">Chen, J. L., Wilson, C. R., and Tapley, B. D.: Satellite gravity measurements confirm accelerated melting of Greenland ice sheet, Science, 313, 1958–1960, 2006.</mixed-citation>
</ref>
<ref id="ref24">
<label>24</label><mixed-citation publication-type="other" xlink:type="simple">Covey, C., Sloan, L. C., and Hoffert, M. I.: Paleoclimate data constraints on climate sensitivity: the paleocalibration method, Climatic Change, 32, 165–184, 1996.</mixed-citation>
</ref>
<ref id="ref25">
<label>25</label><mixed-citation publication-type="other" xlink:type="simple">Cramer, B. S., Toggweiler, J. R., Wright, J. D., Katz, M. E., and Miller, K. G.: Ocean overturning since the Late Cretaceous: inferences from a new benthic foraminiferal isotope compilation, Paleoceanography, 24, PA4216, &lt;a href=&quot;http://dx.doi.org/10.1029/2008PA001683&quot;&gt;https://doi.org/10.1029/2008PA001683&lt;/a&gt;, 2009.</mixed-citation>
</ref>
<ref id="ref26">
<label>26</label><mixed-citation publication-type="other" xlink:type="simple">Crowley, T. J., and Zachos, J. C.: Comparison of zonal temperature profiles for past warm time periods, in: Warm Climates in Earth History, edited by: Huber, B. T., MacLeod, K. G., and Wing, S. L., Cambridge University Press, Cambridge, 50–76, 2000.</mixed-citation>
</ref>
<ref id="ref27">
<label>27</label><mixed-citation publication-type="other" xlink:type="simple">DeConto, R. M. and Pollard, D.: Rapid Cenozoic glaciation of Antarctica induced by declining atmospheric CO&lt;sub&gt;2&lt;/sub&gt;, Nature, 421, 245–249, 2003.</mixed-citation>
</ref>
<ref id="ref28">
<label>28</label><mixed-citation publication-type="other" xlink:type="simple">DeConto, R. M., Pollard, D., Wilson, P. A., Pälike, H., Lear, C. H., and Pagani, M.: Thresholds for Cenozoic bipolar glaciation, Nature, 455, 652–656, 2008.</mixed-citation>
</ref>
<ref id="ref29">
<label>29</label><mixed-citation publication-type="other" xlink:type="simple">Donnadieu, Y., Pierrehumbert, R., Jacob, R., and Fluteau, F.: Modelling the primary control of paleogeography on Cretaceous climate, Earth Planet. Sc. Lett., 248, 426–437, 2006.</mixed-citation>
</ref>
<ref id="ref30">
<label>30</label><mixed-citation publication-type="other" xlink:type="simple">Dumitrescu, M., Brassell, S. C., Schouten, S., Hopmans, E. C., and Sinninghe Damsté, J. S.: Instability in tropical Pacific sea-surface temperatures during the early Aptian, Geology, 34, 833–836, 2006.</mixed-citation>
</ref>
<ref id="ref31">
<label>31</label><mixed-citation publication-type="other" xlink:type="simple">Dunkley Jones, T., Ridgwell, A., Lunt, D. J., Maslin, M. A., Schmidt, D. N., and Valdes, P. J.: A Paleogene perspective on climate sensitivity and methane hydrate instability, Philos. T. Roy. Soc.&amp;nbsp;A, 368, 2395–2415, 2010.</mixed-citation>
</ref>
<ref id="ref32">
<label>32</label><mixed-citation publication-type="other" xlink:type="simple">Dutton, J. E. and Barron, E. J.: Miocene to present vegetation changes: A possible piece of the Cenozoic cooling puzzle, Geology, 25, 39–41, 1997.</mixed-citation>
</ref>
<ref id="ref33">
<label>33</label><mixed-citation publication-type="other" xlink:type="simple">Ellis, T., Hill, P. W., Fenner, N., Williams, G. G., Godbold, D., and Freeman, C.: The interactive effects of elevated carbon dioxide and water table draw-down on carbon cycling in a Welsh ombrotrophic bog, Ecol. Eng., 35, 978–986, 2009.</mixed-citation>
</ref>
<ref id="ref34">
<label>34</label><mixed-citation publication-type="other" xlink:type="simple">Fletcher, B. J., Brentnall, S. J., Anderson, C. W., Berner, R. A., and Beerling, D. J.: Atmospheric carbon dioxide linked with Mesozoic and early Cenozoic climate change, Nat. Geosci., 1, 43–48, 2008.</mixed-citation>
</ref>
<ref id="ref35">
<label>35</label><mixed-citation publication-type="other" xlink:type="simple">Flückiger, J., Blunier, T., Stauffer, B., Chappellaz, J., Spahni, R., Kawamura, K., Schwander, J., Stocker, T. F., and Dahl-Jensen, D.: N&lt;sub&gt;2&lt;/sub&gt;O and CH&lt;sub&gt;4&lt;/sub&gt; variations during the last glacial epoch: Insight into global processes, Global Biogeochem. Cy., 18, GB1020, &lt;a href=&quot;http://dx.doi.org/10.1029/2003GB002122&quot;&gt;https://doi.org/10.1029/2003GB002122&lt;/a&gt;, 2004.</mixed-citation>
</ref>
<ref id="ref36">
<label>36</label><mixed-citation publication-type="other" xlink:type="simple">Forster, A., Schouten, S., Baas, M., and Sinninghe Damsté, J. S.: Mid-Cretaceous (Albian Santonian) sea surface temperature record of the tropical Atlantic Ocean, Geology, 35, 919–922, 2007a.</mixed-citation>
</ref>
<ref id="ref37">
<label>37</label><mixed-citation publication-type="other" xlink:type="simple">Forster, A., Schouten, S., Moriya, K., Wilson, P. A., and Sinninghe Damsté, J. S.: Tropical warming and intermittent cooling during the Cenomanian/Turonian oceanic anoxic event&amp;nbsp;2: sea surface temperature records from the equatorial Atlantic, Paleoceanography, 22, PA1219, &lt;a href=&quot;http://dx.doi.org/10.1029/2006PA001349&quot;&gt;https://doi.org/10.1029/2006PA001349&lt;/a&gt;, 2007b.</mixed-citation>
</ref>
<ref id="ref38">
<label>38</label><mixed-citation publication-type="other" xlink:type="simple">Frakes, L. A., Francis, J. E., and Syktus, J. I.: Climate Modes of the Phanerozoic, Cambridge University Press, Cambridge, 274&amp;nbsp;pp., 1992.</mixed-citation>
</ref>
<ref id="ref39">
<label>39</label><mixed-citation publication-type="other" xlink:type="simple">Freeman, K. H. and Hayes, J. M.: Fractionation of carbon isotopes by phytoplankton and estimates of ancient CO&lt;sub&gt;2&lt;/sub&gt; levels, Global Biogeochem. Cy., 6, 185–198, 1992.</mixed-citation>
</ref>
<ref id="ref40">
<label>40</label><mixed-citation publication-type="other" xlink:type="simple">Gillett, N. P., Arora, V. K., Zickfeld, K., Marshall, S. J., and Merryfield, W. J.: Ongoing climate change following a complete cessation of carbon dioxide emissions, Nat. Geosci., 4, 83–87, 2011.</mixed-citation>
</ref>
<ref id="ref41">
<label>41</label><mixed-citation publication-type="other" xlink:type="simple">Gough, D. O.: Solar interior structure and luminosity variations, Sol. Phys., 74, 21–34, 1981.</mixed-citation>
</ref>
<ref id="ref42">
<label>42</label><mixed-citation publication-type="other" xlink:type="simple">Greenwood, D. R. and Wing, S. L.: Eocene continental climates and latitudinal temperature gradients, Geology, 23, 1044–1048, 1995.</mixed-citation>
</ref>
<ref id="ref43">
<label>43</label><mixed-citation publication-type="other" xlink:type="simple">Greenwood, D. R., Scarr, M. J., and Christophel, D. C.: Leaf stomatal frequency in the Australian tropical rainforest tree &lt;i&gt;Neolitsea dealbata&lt;/i&gt; (Lauraceae) as a proxy measure of atmospheric &lt;i&gt;p&lt;/i&gt;CO&lt;sub&gt;2&lt;/sub&gt;, Palaeogeogr. Palaeocl., 196, 375–393, 2003.</mixed-citation>
</ref>
<ref id="ref44">
<label>44</label><mixed-citation publication-type="other" xlink:type="simple">Hansen, J., Sato, M., Ruedy, R., Nazarenko, L., Lacis, A., Schmidt, G. A., Russell, G., Aleinov, I., Bauer, M., Bauer, S., Bell, N., Cairns, B., Canuto, V., Chandler, M., Cheng, Y., Genio, A. D., Faluvegi, G., Fleming, E., Friend, A., Hall, T., Jackman, C., Kelley, M., Kiang, N., Koch, D., Lean, J., Lerner, J., Lo, K., Menon, S., Miller, R., Minnis, P., Novakov, T., Oinas, V., Perlwitz, J., Perlwitz, J., Rind, D., Romanou, A., Shindell, D., Stone, P., Sun, S., Tausnev, N., Thresher, D., Wielicki, B., Wong, T., Yao, M., and Zhang, S.: Efficacy of climate forcings, J. Geophys. Res., 110, D18104, &lt;a href=&quot;http://dx.doi.org/10.1029/2005JD005776&quot;&gt;https://doi.org/10.1029/2005JD005776&lt;/a&gt;, 2005.</mixed-citation>
</ref>
<ref id="ref45">
<label>45</label><mixed-citation publication-type="other" xlink:type="simple">Hansen, J., Sato, M., Kharecha, P., Beerling, D., Berner, R., Masson-Delmotte, V., Pagani, M., Raymo, M., Royer, D. L., and Zachos, J. C.: Target atmospheric CO&lt;sub&gt;2&lt;/sub&gt;: where should humanity aim?, Open Atmos. Sci. J., 2, 217–231, 2008.</mixed-citation>
</ref>
<ref id="ref46">
<label>46</label><mixed-citation publication-type="other" xlink:type="simple">Haworth, M., Hesselbo, S. P., McElwain, J. C., Robinson, S. A., and Brunt, J. W.: Mid-Cretaceous &lt;i&gt;p&lt;/i&gt;CO&lt;sub&gt;2&lt;/sub&gt; based on stomata of the extinct conifer &lt;i&gt;Pseudofrenelopsis&lt;/i&gt; (Cheirolepidiaceae), Geology, 33, 749–752, 2005.</mixed-citation>
</ref>
<ref id="ref47">
<label>47</label><mixed-citation publication-type="other" xlink:type="simple">Hegerl, G. C., Crowley, T. J., Hyde, W. T., and Frame, D. J.: Climate sensitivity constrained by temperature reconstructions over the past seven centuries, Nature, 440, 1029–1032, 2006.</mixed-citation>
</ref>
<ref id="ref48">
<label>48</label><mixed-citation publication-type="other" xlink:type="simple">Heinemann, M., Jungclaus, J. H., and Marotzke, J.: Warm Paleocene/Eocene climate as simulated in ECHAM5/MPI-OM, Clim. Past, 5, 785–802, &lt;a href=&quot;http://dx.doi.org/10.5194/cp-5-785-2009&quot;&gt;https://doi.org/10.5194/cp-5-785-2009&lt;/a&gt;, 2009.</mixed-citation>
</ref>
<ref id="ref49">
<label>49</label><mixed-citation publication-type="other" xlink:type="simple">Higgins, J. A. and Schrag, D. P.: Beyond methane: Towards a theory for the Paleocene-Eocene Thermal Maximum, Earth Planet. Sc. Lett., 245, 523–537, 2006.</mixed-citation>
</ref>
<ref id="ref50">
<label>50</label><mixed-citation publication-type="other" xlink:type="simple">Hoffert, M. I. and Covey, C.: Deriving global climate sensitivity from palaeoclimate reconstructions, Nature, 360, 573–576, 1992.</mixed-citation>
</ref>
<ref id="ref51">
<label>51</label><mixed-citation publication-type="other" xlink:type="simple">Hollis, C. J., Handley, L., Crouch, E. M., Morgans, H. E. G., Baker, J. A., Creech, J., Collins, K. S., Gibbs, S. J., Huber, M., Schouten, S., Zachos, J. C., and Pancost, R. D.: Tropical sea temperatures in the high-latitude South Pacific during the Eocene, Geology, 37, 99–102, 2009.</mixed-citation>
</ref>
<ref id="ref52">
<label>52</label><mixed-citation publication-type="other" xlink:type="simple">Huber, M.: A hotter greenhouse?, Science, 321, 353–354, 2008.</mixed-citation>
</ref>
<ref id="ref53">
<label>53</label><mixed-citation publication-type="other" xlink:type="simple">Jagniecki, E., Lowenstein, T. K., and Jenkins, D.: Sodium carbonates: temperature and &lt;i&gt;p&lt;/i&gt;CO&lt;sub&gt;2&lt;/sub&gt; indicators for ancient and modern alkaline saline lakes, Geol. Soc. Am. Abstracts with Programs, Paper No.&amp;nbsp;165-402, 42(5), 404, 2010.</mixed-citation>
</ref>
<ref id="ref54">
<label>54</label><mixed-citation publication-type="other" xlink:type="simple">Kiehl, J. T.: Challenges in modeling warm Cenozoic climates, Geochim. Cosmochim. Acta, 73(S1), A648, 2009.</mixed-citation>
</ref>
<ref id="ref55">
<label>55</label><mixed-citation publication-type="other" xlink:type="simple">Kiehl, J. T.: Lessons from Earth&apos;s past, Science, 331, 158–159, 2011.</mixed-citation>
</ref>
<ref id="ref56">
<label>56</label><mixed-citation publication-type="other" xlink:type="simple">Kim, J.-H., Schouten, S., Hopmans, E. C., Donner, B., and Sinninghe Damsté, J. S.: Global sediment core-top calibration of the TEX&lt;sub&gt;86&lt;/sub&gt; paleothermometer in the ocean, Geochim. Cosmochim. Acta, 72, 1154–1173, 2008.</mixed-citation>
</ref>
<ref id="ref57">
<label>57</label><mixed-citation publication-type="other" xlink:type="simple">Kirk-Davidoff, D. B., Schrag, D. P., and Anderson, J. G.: On the feedback of statospheric clouds and polar climate, Geophys. Res. Lett., 29, 1556, &lt;a href=&quot;http://dx.doi.org/10.1029/2002GL014659&quot;&gt;https://doi.org/10.1029/2002GL014659&lt;/a&gt;, 2002.</mixed-citation>
</ref>
<ref id="ref58">
<label>58</label><mixed-citation publication-type="other" xlink:type="simple">Klochko, K., Kaufman, A. J., Yao, W. S., Byrne, R. H., and Tossell, J. A.: Experimental measurement of boron isotope fractionation in seawater, Earth Planet. Sc. Lett., 248, 276–285, 2006.</mixed-citation>
</ref>
<ref id="ref59">
<label>59</label><mixed-citation publication-type="other" xlink:type="simple">Klochko, K., Cody, G. D., Tossell, J. A., Dera, P., and Kaufman, A. J.: Re-evaluating boron speciation in biogenic calcite and aragonite using &lt;sup&gt;11&lt;/sup&gt;B&amp;nbsp;MAS&amp;nbsp;NMR, Geochim. Cosmochim. Acta, 73, 1890–1900, 2009.</mixed-citation>
</ref>
<ref id="ref60">
<label>60</label><mixed-citation publication-type="other" xlink:type="simple">Knutti, R. and Hegerl, G. C.: The equilibrium sensitivity of the Earth&apos;s temperature to radiation changes, Nat. Geosci., 1, 735–743, 2008.</mixed-citation>
</ref>
<ref id="ref61">
<label>61</label><mixed-citation publication-type="other" xlink:type="simple">Kürschner, W. M., Wagner, F., Dilcher, D. L., and Visscher, H.: Using fossil leaves for the reconstruction of Cenozoic paleoatmospheric CO&lt;sub&gt;2&lt;/sub&gt; concentrations, in: Geological Perspectives of Global Climate Change, edited by: Gerhard, L. C., Harrison, W. E., and Hanson, B. M., The American Association of Petroleum Geologists, Tulsa, 169–189, 2001.</mixed-citation>
</ref>
<ref id="ref62">
<label>62</label><mixed-citation publication-type="other" xlink:type="simple">Kump, L. R. and Pollard, D.: Amplification of Cretaceous warmth by biological cloud feedbacks, Science, 320, 195, 2008.</mixed-citation>
</ref>
<ref id="ref63">
<label>63</label><mixed-citation publication-type="other" xlink:type="simple">Lemarchand, D., Gaillardet, J., Lewin, É., and Allègre, C. J.: The influence of rivers on marine boron isotopes and implications for reconstructing past ocean pH, Nature, 408, 951–954, 2000.</mixed-citation>
</ref>
<ref id="ref64">
<label>64</label><mixed-citation publication-type="other" xlink:type="simple">Lloyd, A. H.: Ecological histories from Alaskan tree lines provide insight into future change, Ecology, 86, 1687–1695, 2005.</mixed-citation>
</ref>
<ref id="ref65">
<label>65</label><mixed-citation publication-type="other" xlink:type="simple">Lowenstein, T. K. and Demicco, R. V.: Elevated Eocene atmospheric CO&lt;sub&gt;2&lt;/sub&gt; and its subsequent decline, Science, 313, 1928, 2006.</mixed-citation>
</ref>
<ref id="ref66">
<label>66</label><mixed-citation publication-type="other" xlink:type="simple">Lunt, D. J., Haywood, A. M., Schmidt, G. A., Salzmann, U., Valdes, P. J., and Dowsett, H. J.: Earth system sensitivity inferred from Pliocene modelling and data, Nat. Geosci., 3, 60–64, 2010.</mixed-citation>
</ref>
<ref id="ref67">
<label>67</label><mixed-citation publication-type="other" xlink:type="simple">Matthews, H. D. and Caldeira, K.: Stabilizing climate requires near-zero emissions, Geophys. Res. Lett., 35, L04705, &lt;a href=&quot;http://dx.doi.org/10.1029/2007GL032388&quot;&gt;https://doi.org/10.1029/2007GL032388&lt;/a&gt;, 2008.</mixed-citation>
</ref>
<ref id="ref68">
<label>68</label><mixed-citation publication-type="other" xlink:type="simple">Megonigal, J. P. and Schlesinger, W. H.: Enhanced CH&lt;sub&gt;4&lt;/sub&gt; emission from a wetland soil exposed to elevated CO&lt;sub&gt;2&lt;/sub&gt;, Biogeochemistry, 37, 77–88, 1997.</mixed-citation>
</ref>
<ref id="ref69">
<label>69</label><mixed-citation publication-type="other" xlink:type="simple">Montenegro, A., Brovkin, V., Eby, M., Archer, D., and Weaver, A. J.: Long term fate of anthropogenic carbon, Geophys. Res. Lett., 34, L19707, &lt;a href=&quot;http://dx.doi.org/10.1029/2007GL030905&quot;&gt;https://doi.org/10.1029/2007GL030905&lt;/a&gt;, 2007.</mixed-citation>
</ref>
<ref id="ref70">
<label>70</label><mixed-citation publication-type="other" xlink:type="simple">Moriya, K., Wilson, P. A., Friedrich, O., Erbacher, J., and Kawahata, H.: Testing for ice sheets during the mid-Cretaceous greenhouse using glassy foraminiferal calcite from the mid-Cenomanian tropics on Demerara Rise, Geology, 35, 615–618, 2007.</mixed-citation>
</ref>
<ref id="ref71">
<label>71</label><mixed-citation publication-type="other" xlink:type="simple">Norris, R. D., Bice, K. L., Magno, E. A., and Wilson, P. A.: Jiggling the tropical thermostat in the Cretaceous hothouse, Geology, 30, 299–302, 2002.</mixed-citation>
</ref>
<ref id="ref72">
<label>72</label><mixed-citation publication-type="other" xlink:type="simple">Otto-Bliesner, B. L. and Upchurch, G. R.: Vegetation-induced warming of high-latitude regions during the Late Cretaceous period, Nature, 385, 804–807, 1997.</mixed-citation>
</ref>
<ref id="ref73">
<label>73</label><mixed-citation publication-type="other" xlink:type="simple">Pagani, M., Lemarchand, D., Spivack, A., and Gaillardet, J.: A critical evaluation of the boron isotope-&lt;i&gt;p&lt;/i&gt;H proxy: The accuracy of ancient ocean &lt;i&gt;p&lt;/i&gt;H estimates, Geochim. Cosmochim. Acta, 69, 953–961, 2005.</mixed-citation>
</ref>
<ref id="ref74">
<label>74</label><mixed-citation publication-type="other" xlink:type="simple">Pagani, M., Caldeira, K., Archer, D., and Zachos, J. C.: An ancient carbon mystery, Science, 314, 1556–1557, 2006.</mixed-citation>
</ref>
<ref id="ref75">
<label>75</label><mixed-citation publication-type="other" xlink:type="simple">Pagani, M., Liu, Z., LaRiviere, J., and Ravelo, A. C.: High Earth-system climate sensitivity determined from Pliocene carbon dioxide concentrations, Nat. Geosci., 3, 27–30, 2010.</mixed-citation>
</ref>
<ref id="ref76">
<label>76</label><mixed-citation publication-type="other" xlink:type="simple">Park, J. and Royer, D. L.: Geologic constraints on the glacial amplification of Phanerozoic climate sensitivity, Am. J. Sci., in press, 2011.</mixed-citation>
</ref>
<ref id="ref77">
<label>77</label><mixed-citation publication-type="other" xlink:type="simple">Passalia, M. G.: Cretaceous &lt;i&gt;p&lt;/i&gt;CO&lt;sub&gt;2&lt;/sub&gt; estimation from stomatal frequency analysis of gymnosperm leaves of Patagonia, Argentina, Palaeogeogr. Palaeocl., 273, 17–24, 2009.</mixed-citation>
</ref>
<ref id="ref78">
<label>78</label><mixed-citation publication-type="other" xlink:type="simple">Pearson, P. N. and Palmer, M. R.: Atmospheric carbon dioxide concentrations over the past 60 million years, Nature, 406, 695–699, 2000.</mixed-citation>
</ref>
<ref id="ref79">
<label>79</label><mixed-citation publication-type="other" xlink:type="simple">Pearson, P. N., Ditchfield, P. W., Singano, J., Harcourt-Brown, K. G., Nicholas, C. J., Olsson, R. K., Shackleton, N. J., and Hall, M. A.: Warm tropical sea surface temperatures in the Late Cretaceous and Eocene epochs, Nature, 413, 481–487, 2001.</mixed-citation>
</ref>
<ref id="ref80">
<label>80</label><mixed-citation publication-type="other" xlink:type="simple">Pearson, P. N., van Dongen, B. E., Nicholas, C. J., Pancost, R. D., Schouten, S., Singano, J. M., and Wade, B. S.: Stable warm tropical climate through the Eocene Epoch, Geology, 35, 211–214, 2007.</mixed-citation>
</ref>
<ref id="ref81">
<label>81</label><mixed-citation publication-type="other" xlink:type="simple">Pearson, P. N., Foster, G. L., and Wade, B. S.: Atmospheric carbon dioxide through the Eocene-Oligocene climate transition, Nature, 461, 1110–1113, 2009.</mixed-citation>
</ref>
<ref id="ref82">
<label>82</label><mixed-citation publication-type="other" xlink:type="simple">Pollard, D. and DeConto, R. M.: Hysteresis in Cenozoic Antarctic ice-sheet variations, Global Planet. Change, 45, 9–21, 2005.</mixed-citation>
</ref>
<ref id="ref83">
<label>83</label><mixed-citation publication-type="other" xlink:type="simple">Pucéat, E., Lécuyer, C., Donnadieu, Y., Naveau, P., Cappetta, H., Ramstein, G., Huber, B. T., and Kriwet, J.: Fish tooth δ&lt;sup&gt;18&lt;/sup&gt;O revising Late Cretaceous meridional upper ocean water temperature gradients, Geology, 35, 107–110, 2007.</mixed-citation>
</ref>
<ref id="ref84">
<label>84</label><mixed-citation publication-type="other" xlink:type="simple">Quan, C., Sun, G., and Zhou, Z.: A new Tertiary &lt;i&gt;Ginkgo&lt;/i&gt; (Ginkgoaceae) from the Wuyun Formation of Jiayin, Heilongjiang, northeastern China and its paleoenvironmental implications, Am. J. Bot., 97, 446–457, 2010.</mixed-citation>
</ref>
<ref id="ref85">
<label>85</label><mixed-citation publication-type="other" xlink:type="simple">Raymo, M. E., Grant, B., Horowitz, M., and Rau, G. H.: Mid-Pliocene warmth: stronger greenhouse and stronger conveyor, Mar. Micropaleontol., 27, 313–326, 1996.</mixed-citation>
</ref>
<ref id="ref86">
<label>86</label><mixed-citation publication-type="other" xlink:type="simple">Retallack, G. J.: Greenhouse crises of the past 300 million years, Geol. Soc. Am. Bull., 121, 1441–1455, 2009.</mixed-citation>
</ref>
<ref id="ref87">
<label>87</label><mixed-citation publication-type="other" xlink:type="simple">Roe, G. H. and Baker, M. B.: Why is climate sensitivity so unpredictable?, Science, 318, 629–632, 2007.</mixed-citation>
</ref>
<ref id="ref88">
<label>88</label><mixed-citation publication-type="other" xlink:type="simple">Royer, D. L.: Estimating latest Cretaceous and Tertiary atmospheric CO&lt;sub&gt;2&lt;/sub&gt; concentration from stomatal indices, in: Causes and Consequences of Globally Warm Climates in the Early Paleogene, edited by: Wing, S. L., Gingerich, P. D., Schmitz, B., and Thomas, E., Geological Society of America Special Paper&amp;nbsp;369, Boulder, 79–93, 2003.</mixed-citation>
</ref>
<ref id="ref89">
<label>89</label><mixed-citation publication-type="other" xlink:type="simple">Royer, D. L.: CO&lt;sub&gt;2&lt;/sub&gt;-forced climate thresholds during the Phanerozoic, Geochim. Cosmochim. Acta, 70, 5665–5675, 2006.</mixed-citation>
</ref>
<ref id="ref90">
<label>90</label><mixed-citation publication-type="other" xlink:type="simple">Royer, D. L.: Fossil soils constrain ancient climate sensitivity, P. Natl. Acad. Sci. USA, 107, 517–518, 2010.</mixed-citation>
</ref>
<ref id="ref91">
<label>91</label><mixed-citation publication-type="other" xlink:type="simple">Royer, D. L., Berner, R. A., and Beerling, D. J.: Phanerozoic CO&lt;sub&gt;2&lt;/sub&gt; change: evaluating geochemical and paleobiological approaches, Earth-Sci. Rev., 54, 349–392, 2001a.</mixed-citation>
</ref>
<ref id="ref92">
<label>92</label><mixed-citation publication-type="other" xlink:type="simple">Royer, D. L., Wing, S. L., Beerling, D. J., Jolley, D. W., Koch, P. L., Hickey, L. J., and Berner, R. A.: Paleobotanical evidence for near present-day levels of atmospheric CO&lt;sub&gt;2&lt;/sub&gt; during part of the Tertiary, Science, 292, 2310–2313, 2001b.</mixed-citation>
</ref>
<ref id="ref93">
<label>93</label><mixed-citation publication-type="other" xlink:type="simple">Royer, D. L., Berner, R. A., and Park, J.: Climate sensitivity constrained by CO&lt;sub&gt;2&lt;/sub&gt; concentrations over the past 420 million years, Nature, 446, 530–532, 2007.</mixed-citation>
</ref>
<ref id="ref94">
<label>94</label><mixed-citation publication-type="other" xlink:type="simple">Rustad, J. R. and Zarzycki, P.: Calculation of site-specific carbon-isotope fractionation in pedogenic oxide minerals, P. Natl. Acad. Sci. USA, 105, 10297–10301, 2008.</mixed-citation>
</ref>
<ref id="ref95">
<label>95</label><mixed-citation publication-type="other" xlink:type="simple">Saarnio, S., Saarinen, T., Vasander, H., and Silvola, J.: A moderate increase in the annual CH&lt;sub&gt;4&lt;/sub&gt; efflux by raised CO&lt;sub&gt;2&lt;/sub&gt; or NH&lt;sub&gt;4&lt;/sub&gt;NO&lt;sub&gt;3&lt;/sub&gt; supply in a boreal oligotrophic mire, Global Change Biol., 6, 137–144, 2000.</mixed-citation>
</ref>
<ref id="ref96">
<label>96</label><mixed-citation publication-type="other" xlink:type="simple">Schouten, S., Hopmans, E. C., Forster, A., van Breugel, Y., Kuypers, M. M. M., and Sinninghe Damsté, J. S.: Extremely high sea-surface temperatures at low latitudes during the middle Cretaceous as revealed by archaeal membrane lipids, Geology, 31, 1069–1072, 2003.</mixed-citation>
</ref>
<ref id="ref97">
<label>97</label><mixed-citation publication-type="other" xlink:type="simple">Schrag, D. P. and Alley, R. B.: Ancient lessons for our future climate, Science, 306, 821–822, 2004.</mixed-citation>
</ref>
<ref id="ref98">
<label>98</label><mixed-citation publication-type="other" xlink:type="simple">Seki, O., Foster, G. L., Schmidt, D. N., Mackensen, A., Kawamura, K., and Pancost, R. D.: Alkenone and boron-based Pliocene &lt;i&gt;p&lt;/i&gt;CO&lt;sub&gt;2&lt;/sub&gt; records, Earth Planet. Sc. Lett., 292, 201–211, 2010.</mixed-citation>
</ref>
<ref id="ref99">
<label>99</label><mixed-citation publication-type="other" xlink:type="simple">Sexton, P. F., Wilson, P. A., and Pearson, P. N.: Microstructural and geochemical perspectives on planktic foraminiferal preservation: &quot;glassy&quot; versus &quot;frosty&quot;, Geochem. Geophy. Geosy., 7, Q12P19, &lt;a href=&quot;http://dx.doi.org/10.1029/2006GC001291&quot;&gt;https://doi.org/10.1029/2006GC001291&lt;/a&gt;, 2006.</mixed-citation>
</ref>
<ref id="ref100">
<label>100</label><mixed-citation publication-type="other" xlink:type="simple">Shaffer, G., Olsen, S. M., and Pedersen, J. O. P.: Long-term ocean oxygen depletion in response to carbon dioxide emissions from fossil fuels, Nat. Geosci., 2, 105–109, 2009.</mixed-citation>
</ref>
<ref id="ref101">
<label>101</label><mixed-citation publication-type="other" xlink:type="simple">Shellito, C. J., Sloan, L. C., and Huber, M.: Climate model sensitivity to atmospheric CO&lt;sub&gt;2&lt;/sub&gt; levels in the Early-Middle Paleogene, Palaeogeogr. Palaeocl., 193, 113–123, 2003.</mixed-citation>
</ref>
<ref id="ref102">
<label>102</label><mixed-citation publication-type="other" xlink:type="simple">Shellito, C. J., Lamarque, J.-F., and Sloan, L. C.: Early Eocene Arctic climate sensitivity to &lt;i&gt;p&lt;/i&gt;CO&lt;sub&gt;2&lt;/sub&gt; and basin geography, Geophys. Res. Lett., 36, L09707, &lt;a href=&quot;http://dx.doi.org/10.1029/2009GL037248&quot;&gt;https://doi.org/10.1029/2009GL037248&lt;/a&gt;, 2009.</mixed-citation>
</ref>
<ref id="ref103">
<label>103</label><mixed-citation publication-type="other" xlink:type="simple">Silverman, B. W.: Density Estimation for Statistics and Data Analysis, Chapman &amp; Hall, London, 1986.</mixed-citation>
</ref>
<ref id="ref104">
<label>104</label><mixed-citation publication-type="other" xlink:type="simple">Sloan, L. C. and Rea, D. K.: Atmospheric carbon dioxide and early Eocene climate: A general circulation modeling sensitivity study, Palaeogeogr. Palaeocl., 119, 275–292, 1995.</mixed-citation>
</ref>
<ref id="ref105">
<label>105</label><mixed-citation publication-type="other" xlink:type="simple">Sloan, L. C., Walker, J. C. G., Moore, T. C., Rea, D. K., and Zachos, J. C.: Possible methane-induced polar warming in the early Eocene, Nature, 357, 320–322, 1992.</mixed-citation>
</ref>
<ref id="ref106">
<label>106</label><mixed-citation publication-type="other" xlink:type="simple">Sluijs, A., Bijl, P. K., Schouten, S., Röhl, U., Reichart, G.-J., and Brinkhuis, H.: Southern ocean warming, sea level and hydrological change during the Paleocene-Eocene thermal maximum, Clim. Past, 7, 47–61, &lt;a href=&quot;http://dx.doi.org/10.5194/cp-7-47-2011&quot;&gt;https://doi.org/10.5194/cp-7-47-2011&lt;/a&gt;, 2011.</mixed-citation>
</ref>
<ref id="ref107">
<label>107</label><mixed-citation publication-type="other" xlink:type="simple">Smith, R. Y., Greenwood, D. R., and Basinger, J. F.: Estimating paleoatmospheric &lt;i&gt;p&lt;/i&gt;CO&lt;sub&gt;2&lt;/sub&gt; during the Early Eocene Climatic Optimum from stomatal frequency of &lt;i&gt;Ginkgo&lt;/i&gt;, Okanagan Highlands, British Columbia, Canada, Palaeogeogr. Palaeocl., 293, 120–131, 2010.</mixed-citation>
</ref>
<ref id="ref108">
<label>108</label><mixed-citation publication-type="other" xlink:type="simple">Soden, B. J. and Held, I. M.: An assessment of climate feedbacks in coupled ocean-atmosphere models, J. Climate, 19, 3354–3360, 2006.</mixed-citation>
</ref>
<ref id="ref109">
<label>109</label><mixed-citation publication-type="other" xlink:type="simple">Solomon, S., Plattner, G.-K., Knutti, R., and Friedlingstein, P.: Irreversible climate change due to carbon dioxide emissions, P. Natl. Acad. Sci. USA, 106, 1704–1709, 2009.</mixed-citation>
</ref>
<ref id="ref110">
<label>110</label><mixed-citation publication-type="other" xlink:type="simple">Solomon, S., Daniel, J. S., Sanford, T. J., Murphy, D. M., Plattner, G.-K., Knutti, R., and Friedlingstein, P.: Persistence of climate changes due to a range of greenhouse gases, P. Natl. Acad. Sci. USA, 107, 18354–18359, 2010.</mixed-citation>
</ref>
<ref id="ref111">
<label>111</label><mixed-citation publication-type="other" xlink:type="simple">Spahni, R., Chappellaz, J., Stocker, T. F., Loulergue, L., Hausammann, G., Kawamura, K., Flückiger, J., Schwander, J., Raynaud, D., Masson-Delmotte, V., and Jouzel, J.: Atmospheric methane and nitrous oxide of the Late Pleistocene from Antarctic ice cores, Science, 310, 1317–1321, 2005.</mixed-citation>
</ref>
<ref id="ref112">
<label>112</label><mixed-citation publication-type="other" xlink:type="simple">Sturm, M., Racine, C., and Tape, K.: Increasing shrub abundance in the Arctic, Nature, 411, 546–547, 2001.</mixed-citation>
</ref>
<ref id="ref113">
<label>113</label><mixed-citation publication-type="other" xlink:type="simple">Tipple, B. J., Meyers, S. R., and Pagani, M.: Carbon isotope ratio of Cenozoic CO&lt;sub&gt;2&lt;/sub&gt;: a comparative evaluation of available geochemical proxies, Paleoceanography, 25, PA3202, &lt;a href=&quot;http://dx.doi.org/10.1029/2009PA001851&quot;&gt;https://doi.org/10.1029/2009PA001851&lt;/a&gt;, 2010.</mixed-citation>
</ref>
<ref id="ref114">
<label>114</label><mixed-citation publication-type="other" xlink:type="simple">Tripati, A., Delaney, M. L., Zachos, J. C., Anderson, L. D., Kelly, D. C., and Elderfield, H.: Tropical sea-surface temperature reconstruction for the early Paleogene using Mg/Ca ratios of planktonic foraminifera, Paleoceanography, 18(4), 1101, &lt;a href=&quot;http://dx.doi.org/10.1029/2003PA000937&quot;&gt;https://doi.org/10.1029/2003PA000937&lt;/a&gt;, 2003.</mixed-citation>
</ref>
<ref id="ref115">
<label>115</label><mixed-citation publication-type="other" xlink:type="simple">Valdes, P. J.: Warm climate forcing mechanisms, in: Warm Climates in Earth History, edited by: Huber, B. T., MacLeod, K. G., and Wing, S. L., Cambridge University Press, Cambridge, 3–20, 2000.</mixed-citation>
</ref>
<ref id="ref116">
<label>116</label><mixed-citation publication-type="other" xlink:type="simple">van der Burgh, J., Visscher, H., Dilcher, D. L., and Kürschner, W. M.: Paleoatmospheric signatures in Neogene fossil leaves, Science, 260, 1788–1790, 1993.</mixed-citation>
</ref>
<ref id="ref117">
<label>117</label><mixed-citation publication-type="other" xlink:type="simple">Vaughan, A. P. M.: Climate and geology – a Phanerozoic perspective, in: Deep-Time Perspectives on Climate Change: Marrying the Signal from Computer Models and Biological Proxies, edited by: Williams, M., Haywood, A. M., Gregory, F. J., and Schmidt, D. N., The Geological Society and The Micropalaeontological Society, Special Publications, London, 5–59, 2007.</mixed-citation>
</ref>
<ref id="ref118">
<label>118</label><mixed-citation publication-type="other" xlink:type="simple">Wagner, T., Herrle, J. O., Sinninghe Damsté, J. S., Schouten, S., Stüsser, I., and Hofmann, P.: Rapid warming and salinity changes of Cretaceous surface waters in the subtropical North Atlantic, Geology, 36, 203–206, 2008.</mixed-citation>
</ref>
<ref id="ref119">
<label>119</label><mixed-citation publication-type="other" xlink:type="simple">Wilson, P. A. and Norris, R. D.: Warm tropical ocean surface and global anoxia during the mid-Cretaceous period, Nature, 412, 425–429, 2001.</mixed-citation>
</ref>
<ref id="ref120">
<label>120</label><mixed-citation publication-type="other" xlink:type="simple">Wilson, P. A. and Opdyke, B. N.: Equatorial sea-surface temperatures for the Maastrichtian revealed through remarkable preservation of metastable carbonate, Geology, 24, 555–558, 1996.  Wilson, P. A., Norris, R. D., and Cooper, M. J.: Testing the Cretaceous greenhouse hypothesis using glassy foraminiferal calcite from the core of the Turonian tropics on Demerara Rise, Geology, 30, 607–610, 2002.</mixed-citation>
</ref>
<ref id="ref121">
<label>121</label><mixed-citation publication-type="other" xlink:type="simple">Wing, S. L. and Greenwood, D. R.: Fossils and fossil climate: the case for equable continental interiors in the Eocene, Philos. T. Roy. Soc.&amp;nbsp;B, 341, 243–252, 1993.</mixed-citation>
</ref>
<ref id="ref122">
<label>122</label><mixed-citation publication-type="other" xlink:type="simple">Zachos, J., Pagani, M., Sloan, L., Thomas, E., and Billups, K.: Trends, rhythms, and aberrations in global climate 65 Ma to present, Science, 292, 686–693, 2001.</mixed-citation>
</ref>
<ref id="ref123">
<label>123</label><mixed-citation publication-type="other" xlink:type="simple">Zachos, J. C., Dickens, G. R., and Zeebe, R. E.: An early Cenozoic perspective on greenhouse warming and carbon-cycle dynamics, Nature, 451, 279–283, 2008.</mixed-citation>
</ref>
<ref id="ref124">
<label>124</label><mixed-citation publication-type="other" xlink:type="simple">Zeebe, R. E., Zachos, J. C., and Dickens, G. R.: Carbon dioxide forcing alone insufficient to explain Palaeocene-Eocene Thermal Maximum warming, Nat. Geosci., 2, 576–580, 2009.</mixed-citation>
</ref>
</ref-list>
</back>
</article>