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  <front>
    <journal-meta><journal-id journal-id-type="publisher">ESD</journal-id><journal-title-group>
    <journal-title>Earth System Dynamics</journal-title>
    <abbrev-journal-title abbrev-type="publisher">ESD</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Earth Syst. Dynam.</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">2190-4987</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/esd-17-607-2026</article-id><title-group><article-title>Assessing the performance of LPJmL-5 in simulating vegetation responses to normal and multi-year droughts</article-title><alt-title>Performance of LPJmL-5 in simulating vegetation responses to droughts</alt-title>
      </title-group>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1">
          <name><surname>Ruijsch</surname><given-names>Denise</given-names></name>
          <email>d.ruijsch@uu.nl</email>
        <ext-link>https://orcid.org/0009-0003-9470-2723</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Hauswirth</surname><given-names>Sandra Margrit</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-9208-1757</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>Biemans</surname><given-names>Hester</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-8750-2553</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Wanders</surname><given-names>Niko</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-7102-5454</ext-link></contrib>
        <aff id="aff1"><label>1</label><institution>Department of Physical Geography, Utrecht University, Princetonlaan 8a, Utrecht, the Netherlands</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Water Systems and Global Change, Department of Environmental Sciences, Wageningen University &amp; Research, Droevendaalsesteeg 4, Wageningen, the Netherlands</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Denise Ruijsch (d.ruijsch@uu.nl)</corresp></author-notes><pub-date><day>22</day><month>May</month><year>2026</year></pub-date>
      
      <volume>17</volume>
      <issue>3</issue>
      <fpage>607</fpage><lpage>630</lpage>
      <history>
        <date date-type="received"><day>7</day><month>October</month><year>2025</year></date>
           <date date-type="rev-request"><day>3</day><month>November</month><year>2025</year></date>
           <date date-type="rev-recd"><day>24</day><month>April</month><year>2026</year></date>
           <date date-type="accepted"><day>11</day><month>May</month><year>2026</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2026 Denise Ruijsch et al.</copyright-statement>
        <copyright-year>2026</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026.html">This article is available from https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026.html</self-uri><self-uri xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026.pdf">The full text article is available as a PDF file from https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026.pdf</self-uri>
      <abstract><title>Abstract</title>

      <p id="d2e116">Climate change is expected to increase the frequency and severity of Multi-Year Droughts (MYDs), but their impacts on vegetation remain poorly understood. While satellite records offer valuable insights, they only cover recent decades, limiting the number of MYDs available for analysis. The dynamic global vegetation model LPJmL-5 can simulate vegetation dynamics under varying climate conditions and over longer temporal scales than are typically available from satellite observations. However, its ability to capture vegetation responses to drought, including MYDs, has not yet been systematically evaluated against observation-based datasets. In this study, we benchmark LPJmL-5 against MODIS-derived gross primary production (GPP) to assess how well the model reproduces vegetation responses to drought. We find that LPJmL-5 captures the key temporal and spatial dynamics of drought-related GPP observed by MODIS, although notable differences remain. In particular, LPJmL-5 tends to overestimate vegetation response at the onset of MYDs and shows some rapid recovery behaviour, resulting in muted overall drought impacts. Vegetation responses also vary by type: drought dynamics in croplands are captured relatively well, whereas responses in boreal and temperate vegetation are underestimated in magnitude. These discrepancies appear to be linked to simplified model representations of vegetation stress and mortality, which limit long-term vegetation loss. Beyond drought conditions, LPJmL-5 reproduces absolute GPP values reasonably well in some regions, but performance declines in parts of the Southern Hemisphere and in cropland-dominated areas. This suggests that general GPP simulation performance is not necessarily linked to performance during drought conditions. Overall, this benchmarking highlights strengths and limitations in how LPJmL-5 represents vegetation responses to drought and provides a foundation for future studies of vegetation responses to multi-year droughts.</p>
  </abstract>
    
<funding-group>
<award-group id="gs1">
<funding-source>European Research Council</funding-source>
<award-id>101075354</award-id>
</award-group>
</funding-group>
</article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction</title>
      <p id="d2e128">Multi-Year Droughts (MYDs) can have severe impacts on natural vegetation and croplands, ranging from reduced vegetation health to diminished ecosystem resilience and increased tree mortality <xref ref-type="bibr" rid="bib1.bibx7 bib1.bibx10 bib1.bibx16 bib1.bibx1 bib1.bibx8 bib1.bibx13 bib1.bibx34 bib1.bibx23 bib1.bibx61 bib1.bibx21" id="paren.1"/>. Although less frequent than shorter droughts, MYDs have already been observed worldwide <xref ref-type="bibr" rid="bib1.bibx53" id="paren.2"/> and are projected to increase in both frequency and severity <xref ref-type="bibr" rid="bib1.bibx50" id="paren.3"/>. Understanding how vegetation responds to MYDs is therefore essential for assessing ecosystem vulnerability and anticipating future changes in vegetation health and productivity.</p>
      <p id="d2e140">Satellite remote sensing datasets, such as those derived from MODIS, have been widely used to monitor vegetation dynamics and assess drought impacts <xref ref-type="bibr" rid="bib1.bibx42 bib1.bibx52 bib1.bibx13 bib1.bibx65" id="paren.4"/>. These datasets offer global coverage with high spatial and temporal resolution, making them valuable tools for analysing vegetation responses during droughts. These remotely sensed observations show that, while MYDs generally reduce vegetation productivity and increase stress, the magnitude and direction of these impacts vary across regions. Local water and energy availability can influence responses, resulting in weak, neutral, or even positive effects in some cases <xref ref-type="bibr" rid="bib1.bibx42 bib1.bibx52 bib1.bibx13 bib1.bibx65" id="paren.5"/>. However, satellite records typically extend back only to around 2000 <xref ref-type="bibr" rid="bib1.bibx11 bib1.bibx12" id="paren.6"/>, limiting the number of MYDs available for analysis and reducing the statistical robustness of drought impact assessments based solely on remote sensing datasets.</p>
      <p id="d2e152">Dynamic Global Vegetation Models (DGVMs), such as LPJmL-5 <xref ref-type="bibr" rid="bib1.bibx56" id="paren.7"/>, can simulate vegetation dynamics under varying climate conditions and over longer temporal scales than are typically available from satellite observations. LPJmL-5 has been widely used to study vegetation and agricultural dynamics under historical and projected climate scenarios <xref ref-type="bibr" rid="bib1.bibx56 bib1.bibx35 bib1.bibx15" id="paren.8"/>, and its performance has been extensively evaluated <xref ref-type="bibr" rid="bib1.bibx47 bib1.bibx46 bib1.bibx56" id="paren.9"/>. However, while the model performs well in many aspects, its ability to capture vegetation responses to (multi-year) droughts has not yet been systematically evaluated against observation-based datasets.</p>
      <p id="d2e164">In this study, we benchmark LPJmL-5 against MODIS-derived GPP to evaluate how well the model reproduces vegetation responses to both MYDs and normal droughts (NDs; droughts lasting less than one year). Specifically, we assess the model's ability to capture the magnitude, timing, and spatial patterns of drought-induced GPP anomalies across different regions and vegetation types. This evaluation provides insight into observed drought response behaviour and highlights where LPJmL-5 realistically represents drought stress and recovery, as well as where improvements are needed.</p>

      <fig id="F1" specific-use="star"><label>Figure 1</label><caption><p id="d2e170">Schematic overview of the methodological workflow used in this study. The processes include drought classification, LPJmL-5 model simulations, comparison with MODIS-derived GPP, and the analysis of vegetation responses to both MYDs and NDs across selected focus regions. Blue elements represent observational data, while pink elements indicate model outputs.</p></caption>
        <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f01.png"/>

      </fig>

</sec>
<sec id="Ch1.S2">
  <label>2</label><title>Materials and methods</title>
      <p id="d2e187">This section outlines the definitions, vegetation model, data, and methods used to evaluate vegetation responses to (multi-year) droughts (Fig. <xref ref-type="fig" rid="F1"/>). Section <xref ref-type="sec" rid="Ch1.S2.SS1"/> defines MYDs and NDs, Sect. <xref ref-type="sec" rid="Ch1.S2.SS2"/> describes the LPJmL-5 model, followed by model validation against observational data in Sect. <xref ref-type="sec" rid="Ch1.S2.SS3"/>. Section <xref ref-type="sec" rid="Ch1.S2.SS4"/> and <xref ref-type="sec" rid="Ch1.S2.SS5"/> explain how vegetation responses to (multi-year) droughts are defined. Finally, Sect. <xref ref-type="sec" rid="Ch1.S2.SS6"/> introduces the focus regions analysed in this study.</p>
<sec id="Ch1.S2.SS1">
  <label>2.1</label><title>Drought identification</title>
      <p id="d2e212">Drought indices are essential for studying drought events, as they allow for consistent comparison of drought characteristics across space and time. In this study, we used the Standardized Precipitation Evapotranspiration Index (SPEI), which accounts for both water supply (precipitation, <inline-formula><mml:math id="M1" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula>) and atmospheric water demand (potential evapotranspiration, PET) <xref ref-type="bibr" rid="bib1.bibx54" id="paren.10"/>. SPEI is particularly suitable for assessing drought impacts on vegetation as vegetation is not only impacted by lack of water supply via <inline-formula><mml:math id="M2" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula>, but also the increase in atmospheric water demand via PET.</p>
      <p id="d2e232">SPEI was computed at multiple timescales (1–24 months) using <inline-formula><mml:math id="M3" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula> and PET derived from W5E5 reanalysis data (see Table <xref ref-type="table" rid="T1"/>, <xref ref-type="bibr" rid="bib1.bibx28" id="altparen.11"/>). Both <inline-formula><mml:math id="M4" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula> and PET were aggregated to monthly values to generate monthly SPEI outputs. A detailed description of the SPEI calculation is provided in Sect. S1 in the Supplement.</p>

<table-wrap id="T1" specific-use="star"><label>Table 1</label><caption><p id="d2e257">Overview of datasets and variables used in this study. W5E5 climate variables are grouped by their application in either SPEI calculation or LPJmL-5 simulations.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="left"/>
     <oasis:colspec colnum="4" colname="col4" align="left"/>
     <oasis:colspec colnum="5" colname="col5" align="justify" colwidth="3.5cm"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Dataset/Variable</oasis:entry>
         <oasis:entry colname="col2">Temporal range</oasis:entry>
         <oasis:entry colname="col3">Temporal resolution</oasis:entry>
         <oasis:entry colname="col4">Spatial resolution</oasis:entry>
         <oasis:entry colname="col5" align="left">Use/Reference</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row rowsep="1">
         <oasis:entry namest="col1" nameend="col5">W5E5 Climate Forcing Data </oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Used for SPEI calculation</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx28" id="text.12"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Precipitation [mm d<sup>−1</sup>]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5° (<inline-formula><mml:math id="M6" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">55</mml:mn></mml:mrow></mml:math></inline-formula> km)</oasis:entry>
         <oasis:entry colname="col5" align="left">SPEI input</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Temperature [°C]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">PET calc. for SPEI</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Relative humidity [%]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">PET calc. for SPEI</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Wind speed [m s<sup>−1</sup>]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">PET calc. for SPEI</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Surface pressure [Pa]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">PET calc. for SPEI</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Downward shortwave radiation [W m<sup>−2</sup>]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">PET calc. for SPEI</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Used in LPJmL-5 simulations</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx28" id="text.13"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Precipitation [mm d<sup>−1</sup>]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">LPJmL-5 input</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Temperature [°C]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">LPJmL-5 input</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Wind speed [m s<sup>−1</sup>]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">LPJmL-5 input</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Downward longwave radiation [W m<sup>−2</sup>]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">LPJmL-5 input</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Downward shortwave radiation [W m<sup>−2</sup>]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Daily</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">LPJmL-5 input</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry namest="col1" nameend="col5">Additional LPJmL-5 Inputs </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">CO<sub>2</sub> concentration [ppm]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Annual</oasis:entry>
         <oasis:entry colname="col4">Global</oasis:entry>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx4" id="text.14"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">NH<sub><italic>x</italic></sub> deposition [g m<sup>−2</sup> per month]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Monthly</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx63" id="text.15"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">NO<sub><italic>y</italic></sub> deposition [g m<sup>−2</sup> per month]</oasis:entry>
         <oasis:entry colname="col2">1901–2019</oasis:entry>
         <oasis:entry colname="col3">Monthly</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx63" id="text.16"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Soil pH</oasis:entry>
         <oasis:entry colname="col2">–</oasis:entry>
         <oasis:entry colname="col3">–</oasis:entry>
         <oasis:entry colname="col4">0.5°</oasis:entry>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx55" id="text.17"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Soil type</oasis:entry>
         <oasis:entry colname="col2">–</oasis:entry>
         <oasis:entry colname="col3">–</oasis:entry>
         <oasis:entry colname="col4">250 m (0.25 km)</oasis:entry>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx24" id="text.18"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Land use</oasis:entry>
         <oasis:entry colname="col2">2020</oasis:entry>
         <oasis:entry colname="col3">–</oasis:entry>
         <oasis:entry colname="col4">5 min (<inline-formula><mml:math id="M18" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">9.25</mml:mn></mml:mrow></mml:math></inline-formula> km)</oasis:entry>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx22" id="text.19"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Land cover</oasis:entry>
         <oasis:entry colname="col2">2019</oasis:entry>
         <oasis:entry colname="col3">–</oasis:entry>
         <oasis:entry colname="col4">300 m (0.3 km)</oasis:entry>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx18" id="text.20"/>
                    </oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry namest="col1" nameend="col5">Remote Sensing Datasets </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">MODIS GPP</oasis:entry>
         <oasis:entry colname="col2">2000–2019</oasis:entry>
         <oasis:entry colname="col3">16 d</oasis:entry>
         <oasis:entry colname="col4">1 km</oasis:entry>
         <oasis:entry colname="col5" align="left">
                      <xref ref-type="bibr" rid="bib1.bibx11 bib1.bibx12" id="text.21"/>
                    </oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d2e880">To define MYDs and NDs, we focused on SPEI calculated at a 12-month timescale (SPEI-12), as it captures multi-year drought conditions while minimizing seasonal variability. Following <xref ref-type="bibr" rid="bib1.bibx59" id="text.22"/>, MYDs are defined as periods where SPEI-12 remains below <inline-formula><mml:math id="M19" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula> for at least 12 consecutive months. The threshold of <inline-formula><mml:math id="M20" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula> was chosen to represent moderate to extreme droughts <xref ref-type="bibr" rid="bib1.bibx32" id="paren.23"/>. While the start date of the drought is only identified as the first month where SPEI-12 dips below <inline-formula><mml:math id="M21" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>, this definition already takes into account the accumulated <inline-formula><mml:math id="M22" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula> and PET over the 12 months leading up to the start of the drought. <xref ref-type="bibr" rid="bib1.bibx42" id="text.24"/> evaluated the impact of different SPEI aggregation periods and concluded that the SPEI-12 is suitable for capturing multi-year drought dynamics.</p>
      <p id="d2e930">To distinguish MYDs from shorter drought events, we define NDs as periods where SPEI-12 remains below <inline-formula><mml:math id="M23" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula> for less than 12 consecutive months. This definition ensures that each drought event is consistently categorized as either an MYD or ND, allowing us to systematically compare their respective impacts on vegetation.</p>
</sec>
<sec id="Ch1.S2.SS2">
  <label>2.2</label><title>Dynamic global vegetation modelling using LPJmL-5</title>
      <p id="d2e951">In this study, the DGVM LPJmL-5 (v5.7.9, <xref ref-type="bibr" rid="bib1.bibx56" id="altparen.25"/>) was used to simulate vegetation responses to MYDs. LPJmL-5 builds on LPJmL-4 and simulates both natural and agricultural vegetation, linking their growth and productivity through consistent water, carbon, and energy fluxes <xref ref-type="bibr" rid="bib1.bibx47" id="paren.26"/>. LPJmL-5 makes use of the big-leaf representation for different Plant Functional Types (PFTs) that can coexist within a grid cell and compete for resources such as light, water, and nitrogen <xref ref-type="bibr" rid="bib1.bibx56" id="paren.27"/>. Land cover can either be simulated dynamically or be prescribed, both of which are used in this study. In contrast, Crop Functional Types (CFTs) are always prescribed, with a distinction between irrigated and rainfed crops. Each grid cell can contain both managed and natural vegetation fractions, allowing for a realistic subgrid representation of land-use. CFTs only appear when crops are sown and grow on soil areas separate from natural vegetation <xref ref-type="bibr" rid="bib1.bibx56" id="paren.28"/>. The model also includes key physiological processes such as photosynthesis, gross primary production, evapotranspiration, and plant responses to soil moisture and drought stress <xref ref-type="bibr" rid="bib1.bibx47" id="paren.29"/>. The terrestrial nitrogen cycle is incorporated to account for nutrient limitations to plant growth <xref ref-type="bibr" rid="bib1.bibx56" id="paren.30"/>. This makes that LPJmL-5 contains all the necessary processes to study vegetation responses to drought, though its performance may still vary across regions and vegetation types.</p>
<sec id="Ch1.S2.SS2.SSS1">
  <label>2.2.1</label><title>Drought in the LPJmL-5 model</title>
      <p id="d2e980">In LPJmL-5, GPP is first calculated as a potential GPP, based on the light-limited (JE) and Rubisco-limited (JC) rates and on the absorbed photosynthetically active radiation (APAR; <xref ref-type="bibr" rid="bib1.bibx47" id="altparen.31"/>). This potential GPP is then constrained by soil water availability. When the water supply cannot meet atmospheric demand, water stress develops and the potential GPP is reduced to the actual GPP.</p>
      <p id="d2e986">Drought can further influence GPP through phenology. LPJmL-5 includes a phenology function that controls how active the canopy is, ranging from 0 (no active canopy) to 1 (fully active canopy). When phenology declines, leaf area and FAPAR decrease, thereby reducing GPP. The phenology is determined by four limiting functions: cold temperature, light availability, water availability, and heat stress <xref ref-type="bibr" rid="bib1.bibx47" id="paren.32"/>. Each factor varies between 0 (full limitation) and 1 (no limitation), and their combined effect defines the total phenology. The limiting functions follow logistic curves and include a memory term, meaning that canopy responses are delayed and differ across PFTs <xref ref-type="bibr" rid="bib1.bibx47" id="paren.33"/>. The associated parameters were calibrated by fitting simulated FAPAR to satellite-derived FAPAR over 30 years <xref ref-type="bibr" rid="bib1.bibx47" id="paren.34"/>.</p>
      <p id="d2e998">Vegetation in LPJmL-5 is represented using the big-leaf representation, meaning that all leaves of a PFT within a grid cell are treated as a single canopy layer with an average temperature, photosynthesis, and water use. Under drought, reductions in phenology and water supply therefore shrink this “big leaf” and makes it less productive, rather than simulating individual trees.</p>
      <p id="d2e1001">Finally, LPJmL-5 does not simulate explicit drought-induced mortality. Instead, mortality emerges indirectly from low productivity and heat stress, and is capped by a maximum annual mortality rate <xref ref-type="bibr" rid="bib1.bibx47" id="paren.35"/>. As a result, prolonged water deficits only lead to die-off when plants repeatedly fail to maintain sufficient biomass. Moreover, this mortality process is only active when land cover is simulated dynamically and is not applied when land cover is prescribed.</p>
</sec>
<sec id="Ch1.S2.SS2.SSS2">
  <label>2.2.2</label><title>LPJmL-5 input and output</title>
      <p id="d2e1015">In this study, LPJmL-5 was used to simulate vegetation dynamics at a 0.1° spatial resolution (<inline-formula><mml:math id="M24" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">11</mml:mn></mml:mrow></mml:math></inline-formula> km) and a daily temporal resolution. The model was forced with meteorological forcing, soil properties, land use, and land cover derived from observational datasets (Table <xref ref-type="table" rid="T1"/>). Daily meteorological forcings are taken from the W5E5 dataset and include temperature, precipitation, downward longwave and shortwave radiation, and wind speed <xref ref-type="bibr" rid="bib1.bibx28" id="paren.36"/>. Soil inputs include static variables such as soil pH <xref ref-type="bibr" rid="bib1.bibx55" id="paren.37"/> and soil type <xref ref-type="bibr" rid="bib1.bibx24" id="paren.38"/>, as well as dynamic inputs like monthly nitrogen deposition (NH<sub>4</sub> and NO<sub>3</sub>) <xref ref-type="bibr" rid="bib1.bibx63" id="paren.39"/>. Annual atmospheric CO<sub>2</sub> concentrations were taken from <xref ref-type="bibr" rid="bib1.bibx4" id="text.40"/>. Land use and country code data are provided at a 5 min (<inline-formula><mml:math id="M28" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">9.25</mml:mn></mml:mrow></mml:math></inline-formula> km) spatial resolution, and soil type data at 250 m. All three were regridded to a 0.1° spatial resolution, whereas soil pH and climate forcing data are downscaled using a nearest neighbour interpolation from their original 0.5° (<inline-formula><mml:math id="M29" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">55</mml:mn></mml:mrow></mml:math></inline-formula> km) resolution.</p>
      <p id="d2e1094">The land use map was primarily derived from <xref ref-type="bibr" rid="bib1.bibx22" id="text.41"/>, which provides information on crop areas. However, the crop types in this dataset did not directly match the CFTs used in LPJmL-5, so they were reclassified accordingly. Additionally, pasture areas are a land-use type in LPJmL-5 but were not included in the IFPRI dataset. These were obtained from the ESA land cover maps <xref ref-type="bibr" rid="bib1.bibx18" id="paren.42"/> and incorporated into the land-use input. For the land cover input, we used the ESA PFT maps <xref ref-type="bibr" rid="bib1.bibx18" id="paren.43"/>, which provide fractional cover of tree, shrub, and grass types. Because LPJmL-5 separates PFTs by climate zone (tropical, temperate, and boreal), we reclassified the ESA PFT maps to match the PFTs in LPJmL-5 using the Köppen-Geiger classification <xref ref-type="bibr" rid="bib1.bibx2" id="paren.44"/>, following the approach of <xref ref-type="bibr" rid="bib1.bibx14" id="text.45"/> (see Sect. S6). Shrubs were merged with the corresponding tree PFTs because LPJmL-5 does not distinguish between these growth forms. This procedure ensured that land-use and land-cover inputs were compatible with the vegetation types represented in LPJmL-5.</p>
      <p id="d2e1112">LPJmL-5 provides a multitude of vegetation-related outputs on ecological, hydrological, and agricultural components (such as leaf area index, soil moisture, runoff, and crop yields). To study vegetation response to drought, this study used the monthly Gross Primary Production (GPP), which is the amount of carbon captured from the atmosphere through vegetation photosynthesis <xref ref-type="bibr" rid="bib1.bibx3" id="paren.46"/>. The GPP output used in this study was produced at a 0.1° spatial and monthly temporal resolution.</p>
</sec>
</sec>
<sec id="Ch1.S2.SS3">
  <label>2.3</label><title>Model validation</title>
<sec id="Ch1.S2.SS3.SSS1">
  <label>2.3.1</label><title>MODIS GPP as validation dataset</title>
      <p id="d2e1134">In this study, we evaluated simulated GPP by LPJmL5 against MODIS satellite derived GPP estimates. Although MODIS GPP is widely used, it is not purely observational, but derived using the MOD17 light-use efficiency algorithm and relies on inputs such as MODIS FPAR, meteorological drivers, and land-cover classification <xref ref-type="bibr" rid="bib1.bibx43" id="paren.47"/>. Therefore, before using it as a validation dataset, we assessed how well the MODIS GPP product represents vegetation productivity (Sect. S3).</p>
      <p id="d2e1140">We compared MODIS GPP with two satellite products that are more directly observation-based: the Enhanced Vegetation Index (EVI; <xref ref-type="bibr" rid="bib1.bibx11 bib1.bibx12" id="altparen.48"/>) and solar-induced fluorescence (SIF; <xref ref-type="bibr" rid="bib1.bibx58" id="altparen.49"/>). Both comparisons showed strong and statistically significant relationships, particularly outside tropical regions, indicating that MODIS GPP captures large-scale variability in vegetation productivity. Other validation studies report similar limitations, with reduced performance mainly in tropical ecosystems. A full description of these analyses is provided in Sect. S3. Together, these results support the use of MODIS GPP as a validation product for LPJmL5, while acknowledging that uncertainties are larger in tropical regions.</p>
      <p id="d2e1149">To ensure consistency with LPJmL5 output, the MODIS GPP data was regridded to a spatial resolution of 0.1° and a monthly temporal resolution. Because LPJmL-5 provides a single GPP value per grid cell, each cell was assigned to its dominant vegetation type. CFT fractions were aggregated into one cropland category, while natural PFTs were grouped into broader classes: tropical trees, temperate trees, boreal trees, tropical C<sub>4</sub> grass, temperate C<sub>3</sub> grass, and polar C<sub>3</sub> grass (Appendix A).</p>
</sec>
<sec id="Ch1.S2.SS3.SSS2">
  <label>2.3.2</label><title>Kling-Gupta Efficiency</title>
      <p id="d2e1187">To quantify model performance, we used the Kling-Gupta Efficiency <xref ref-type="bibr" rid="bib1.bibx17" id="paren.50"/>, which combines correlation, bias, and variability into a single metric:

              <disp-formula id="Ch1.E1" content-type="numbered"><label>1</label><mml:math id="M33" display="block"><mml:mrow><mml:mtext>KGE</mml:mtext><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msqrt><mml:mrow><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">α</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">β</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup></mml:mrow></mml:msqrt><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>

            where <inline-formula><mml:math id="M34" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula> is the linear correlation between observations and simulations, <inline-formula><mml:math id="M35" display="inline"><mml:mrow><mml:mi mathvariant="italic">α</mml:mi><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi mathvariant="italic">σ</mml:mi><mml:mi mathvariant="normal">sim</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">σ</mml:mi><mml:mi mathvariant="normal">obs</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:math></inline-formula> a measure of the variability error, and <inline-formula><mml:math id="M36" display="inline"><mml:mrow><mml:mi mathvariant="italic">β</mml:mi><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mi mathvariant="normal">sim</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mi mathvariant="normal">obs</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:math></inline-formula> a bias term. Here, <inline-formula><mml:math id="M37" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">σ</mml:mi><mml:mi mathvariant="normal">obs</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M38" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">σ</mml:mi><mml:mi mathvariant="normal">sim</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are the standard deviations of the observations and simulations, respectively, while <inline-formula><mml:math id="M39" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mi mathvariant="normal">sim</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M40" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mi mathvariant="normal">obs</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are the corresponding means. KGE values greater than <inline-formula><mml:math id="M41" display="inline"><mml:mrow><mml:mi mathvariant="normal">−</mml:mi><mml:mn mathvariant="normal">0.41</mml:mn></mml:mrow></mml:math></inline-formula> indicate that the model performs better than the mean benchmark, while a value of 1 indicates perfect agreement between simulations and observations <xref ref-type="bibr" rid="bib1.bibx25" id="paren.51"/>. This allows us to assess how well LPJmL-5 simulates GPP for different PFTs compared to MODIS, and to identify structural patterns in performance across the globe.</p>
</sec>
</sec>
<sec id="Ch1.S2.SS4">
  <label>2.4</label><title>Vegetation sensitivity to varying drought timescales</title>
      <p id="d2e1378">After evaluating how well LPJmL-5 simulates general patterns of GPP compared to MODIS GPP, we examined how well LPJmL-5 can model vegetation responses to drought. Here, we aim to understand modelled vegetation sensitivity to drought without differentiating between “normal” and “multi-year” droughts. To ensure a fair comparison of vegetation responses across regions and to remove seasonal variation, the GPP data was normalized and standardized using <inline-formula><mml:math id="M42" display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula>-score normalization resulting in a standardized GPP anomaly (GPP<sub>SA</sub>). The reference normalization period was chosen to match the MODIS dataset (2000–2019) to avoid errors resulting from GPP trend differences between modelled and observed data.</p>
      <p id="d2e1397">Vegetation can respond differently to drought events of different durations. To capture this, we used the extreme-based method developed by <xref ref-type="bibr" rid="bib1.bibx9" id="text.52"/>. This method examines how different drought durations affect vegetation across various SPEI timescales (ranging from 1 to 24 months). Specifically, we calculated the average GPP<sub>SA</sub> response for each timescale and identified which one has the strongest negative impact during drought periods. This timescale is referred to as the dominant drought timescale and reflects how quickly vegetation responds to drought stress. Shorter timescales indicate faster, more sensitive responses, while longer timescales suggest a slower response and less drought-sensitive vegetation. For this analysis, a SPEI threshold of <inline-formula><mml:math id="M45" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula> was chosen to define atmospheric drought conditions, as suggested by <xref ref-type="bibr" rid="bib1.bibx32" id="text.53"/>.</p>
</sec>
<sec id="Ch1.S2.SS5">
  <label>2.5</label><title>Vegetation response to MYDs</title>
      <p id="d2e1433">After determining how sensitive vegetation is to drought at different timescales, we further investigated how vegetation in LPJmL-5 responds to MYDs specifically. We compared MODIS observations and LPJmL-5 simulations by analysing global spatial patterns together with temporal dynamics in selected focus regions. Vegetation responses during MYDs were quantified by using the mean GPP<sub>SA</sub> during MYD events.</p>
</sec>
<sec id="Ch1.S2.SS6">
  <label>2.6</label><title>Selection of focus regions</title>
      <p id="d2e1453">While this study primarily examines global drought patterns, it also focuses on selected focus regions to better understand local drought dynamics and how regional differences affect vegetation responses. The focus regions used in this study are: California (CAL), the Rhine-Meuse delta in western Europe (WEU), the Brahmaputra River basin spanning Bangladesh, India, Bhutan, and China (BRA), central Argentina (ARG), the Orange River basin in southern Africa (SA), and the Murray-Darling basin in Australia (AUS) (Appendix B). These regions correspond to the focus regions used by <xref ref-type="bibr" rid="bib1.bibx42" id="text.54"/> examining MYD patterns. The focus regions were selected based on the number of MYDs that have occurred between 2000–2019, geographic and climatic diversity, and variation in vegetation types, ensuring a broad representation of both water- and energy-limited regions across different climate zones.</p>
</sec>
</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Results</title>
      <p id="d2e1469">The following sections describe different aspects of the analysis on the potential of LPJmL-5 to simulate vegetation response to (multi-year) droughts. Section <xref ref-type="sec" rid="Ch1.S3.SS1"/> examines LPJmL's ability to reproduce satellite-derived GPP, while Sect. <xref ref-type="sec" rid="Ch1.S3.SS2"/> and <xref ref-type="sec" rid="Ch1.S3.SS3"/> focus on its ability to reproduce (multi-year) droughts.</p>
<sec id="Ch1.S3.SS1">
  <label>3.1</label><title>General performance of LPJmL-5</title>
      <p id="d2e1485">To evaluate LPJmL-5's ability to simulate GPP, we compared modelled GPP to MODIS-derived GPP using two simulation setups: dynamic and prescribed land cover. The dynamic land cover setup allows vegetation fractions to shift in response to climate and includes background mortality, capturing potential vegetation loss <xref ref-type="bibr" rid="bib1.bibx47" id="paren.55"/>. However, dynamic vegetation does not necessarily match observed vegetation types, which can lead to differences in GPP due to mismatched vegetation types. The prescribed land cover simulation maintains fixed vegetation fractions based on observed maps. This approach ensures that variations in GPP result from the model's ability to simulate vegetation productivity. However, this setup does not include background mortality, potentially causing an underestimation of vegetation loss. By comparing both setups, we can determine which one provides a better representation of satellite-derived GPP and choose the setup to be used in later analyses. Model performance is evaluated by comparing LPJmL-5 and MODIS GPP using the KGE metric (Sect. <xref ref-type="sec" rid="Ch1.S3.SS1.SSS1"/>) and time series analysis (Sect. <xref ref-type="sec" rid="Ch1.S3.SS1.SSS2"/>).</p>

      <fig id="F2" specific-use="star"><label>Figure 2</label><caption><p id="d2e1497">Spatial comparison of LPJmL-5 model performance using the Kling-Gupta Efficiency (KGE) metric and its components for dynamic (left column) and prescribed (right column) land cover simulations. The rows show <bold>(a–b)</bold> overall KGE, followed by its components: <bold>(c–d)</bold> correlation, <bold>(e–f)</bold> bias, and <bold>(g–h)</bold> variability.</p></caption>
          <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f02.png"/>

        </fig>

<sec id="Ch1.S3.SS1.SSS1">
  <label>3.1.1</label><title>Evaluating model performance with the KGE metric</title>
      <p id="d2e1525">Figure <xref ref-type="fig" rid="F2"/> shows the spatial distribution of the KGE and its components (correlation, bias, and variability) for both dynamic and prescribed land cover simulations. Overall, the prescribed land cover demonstrates a clear performance improvement over the dynamic simulation (Fig. <xref ref-type="fig" rid="F2"/>a and b).</p>
      <p id="d2e1532">The dynamic land cover consistently shows low agreement with MODIS across all KGE components (Fig. <xref ref-type="fig" rid="F2"/>c, e and g). The agreement is particularly poor in the Southern Hemisphere, with KGE values often falling below <inline-formula><mml:math id="M47" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.41</mml:mn></mml:mrow></mml:math></inline-formula>, which is the threshold for a skilful performance. In contrast, the Northern Hemisphere generally shows a better performance. Switching to prescribed land cover results in improved KGE scores globally. However, the performance difference between the Northern and Southern Hemispheres remains.</p>
      <p id="d2e1547">When examining the individual KGE components, the correlation between modelled and satellite-derived GPP is relatively high for both simulations, except in the tropical and semi-arid regions (Fig. <xref ref-type="fig" rid="F2"/>c and d). This suggests that the intra-annual response to changing meteorological conditions is similar between LPJmL-5 and satellite-derived MODIS GPP.</p>
      <p id="d2e1552">Both the bias (Fig. <xref ref-type="fig" rid="F2"/>e) and variability (Fig. <xref ref-type="fig" rid="F2"/>g) components are overestimated in the dynamic land cover simulation. The bias frequently exceeds 2, except in the tropical regions, suggesting that the model's mean GPP is approximately double that of the satellite-derived value. The prescribed land cover simulation (Fig. <xref ref-type="fig" rid="F2"/>f) significantly reduces this bias, particularly in tropical and temperate regions, bringing modelled GPP closer to the satellite-derived reference. A similar pattern is observed for the GPP variability, where the dynamic land cover shows overestimated variability (greater than 2) in Fig. <xref ref-type="fig" rid="F2"/>g, while prescribed land cover reduces this variability (Fig. <xref ref-type="fig" rid="F2"/>h).</p>
      <p id="d2e1566">These differences in model performance can be attributed to discrepancies between the modelled and observed vegetation types when using dynamic land cover. For example, we observe that LPJmL-5 simulates tree cover in regions where the ESA land cover map shows predominantly grasslands, such as Western Europe and parts of South Africa (see Appendix C and Sect. S5). These discrepancies in vegetation types lead to differences in GPP that are not related to the model's capability to simulate GPP responses to changing meteorological conditions, but rather to inconsistencies in vegetation cover. By using prescribed land cover fractions based on the ESA land cover map, we observe an improvement in model performance, with remaining errors linked to LPJmL-5's inability to accurately simulate the GPP response to changes in meteorological conditions.</p>

      <fig id="F3" specific-use="star"><label>Figure 3</label><caption><p id="d2e1571">Cumulative distribution function (CDF) of the KGE of the prescribed land cover GPP for the different vegetation types in LPJmL-5. Percentages show the percentage of grid cells with that dominant land cover. Black dashed line indicates KGE <inline-formula><mml:math id="M48" display="inline"><mml:mrow><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.41</mml:mn></mml:mrow></mml:math></inline-formula>. Values greater than <inline-formula><mml:math id="M49" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.41</mml:mn></mml:mrow></mml:math></inline-formula> indicate that the model improved upon the mean benchmark.</p></caption>
            <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f03.png"/>

          </fig>

      <p id="d2e1602">To further assess model performance in the prescribed land cover simulation, we analyse the KGE distribution for each PFT, as shown in Fig. <xref ref-type="fig" rid="F3"/>. This cumulative distribution function (CDF) displays the proportion of grid cells for each dominant vegetation type (Appendix A) that exceed the KGE threshold of <inline-formula><mml:math id="M50" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.41</mml:mn></mml:mrow></mml:math></inline-formula>, which represents the mean benchmark <xref ref-type="bibr" rid="bib1.bibx25" id="paren.56"/>.</p>
      <p id="d2e1620">The performance of LPJmL-5 varies significantly across PFTs. Croplands show the lowest agreement with satellite-derived GPP, with over 60 % of grid cells falling below the KGE <inline-formula><mml:math id="M51" display="inline"><mml:mrow><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.41</mml:mn></mml:mrow></mml:math></inline-formula> line. In contrast, temperate trees show the best agreement, with 90 % showing skilful GPP simulations. Approximately 80 % of areas dominated by boreal trees show good agreement with MODIS data. Polar and temperate C<sub>3</sub> grasses are the dominant PFTs and exhibit intermediate performance, with around 75 % of their area showing significant skill. Tropical trees show moderate KGE values, indicating a moderate performance in simulating the GPP. The corresponding analysis for the dynamic land-cover simulation (Fig. <xref ref-type="fig" rid="FC2"/>) shows consistently lower agreement across most PFTs than for prescribed land cover.</p>
      <p id="d2e1646">Overall, the KGE map and the CDF analysis (Figs. <xref ref-type="fig" rid="F2"/>, <xref ref-type="fig" rid="F3"/>, and <xref ref-type="fig" rid="FC2"/>) indicate that simulating with prescribed land cover outperforms dynamic land cover in LPJmL-5. However, significant improvements in modelled GPP response can still be made in the Southern Hemisphere and for croplands. Based on these results, all subsequent simulations will be done using prescribed land cover to ensure a more robust comparison with observational data.</p>

      <fig id="F4" specific-use="star"><label>Figure 4</label><caption><p id="d2e1658">Time series of the GPP for LPJmL-5 with prescribed land cover (green) and MODIS (black) for the focus regions: California (CAL), western Europe (WEU), Brahmaputra (BRA), middle Argentina (ARG), southern Africa (SA) and the Murray Darling Basin in Australia (AUS). SPEI-12 is shaded in the background, where red areas indicate drier periods and blue areas wetter periods. Additionally, the figure includes the correlation between MODIS and LPJmL-5 per region. Note that WEU uses a different <inline-formula><mml:math id="M53" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> axis range from the other regions.</p></caption>
            <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f04.png"/>

          </fig>

</sec>
<sec id="Ch1.S3.SS1.SSS2">
  <label>3.1.2</label><title>Evaluating model performance with the GPP time series</title>
      <p id="d2e1683">Figure <xref ref-type="fig" rid="F4"/> shows the monthly GPP for LPJmL-5 (with prescribed land cover) and MODIS across the six focus regions between 2000 and 2019. The background shading represents the SPEI-12, highlighting dry periods in red and wet periods in blue. Additionally, the figure includes the correlation between MODIS and LPJmL-5.</p>
      <p id="d2e1688">Three general patterns can be identified in the time series: (1) A temporal shift in the LPJmL-5 GPP time series, with peaks occurring earlier in the season compared to MODIS; (2) Overestimation of summer GPP peaks, and in some cases underestimation of winter values; (3) Lower model performance during dry periods.</p>
      <p id="d2e1691">These patterns are reflected to varying degrees across the focus regions. CAL and BRA show the temporal shift, which may be the cause of the lower correlation with MODIS. BRA and WEU exhibit overestimation of summer GPP peaks, with SA also showing the underestimation of winter values. ARG performs relatively well, although it also shows slightly increased summer peaks. In AUS, the contrast between wet and dry periods is largest, with GPP simulated accurately during wet periods, while the model does not capture dry periods well. Despite these issues, WEU, ARG, SA, and AUS show relatively high correlation values, indicating that LPJmL-5 generally captures the seasonal and interannual patterns of satellite-derived GPP in these regions. Overall, based on the KGE values and the time series analysis, we can conclude that LPJmL-5 simulates GPP reasonably well, although notable regional differences remain.</p>
</sec>
</sec>
<sec id="Ch1.S3.SS2">
  <label>3.2</label><title>General drought response</title>
      <p id="d2e1703">To evaluate the ability of LPJmL-5 to capture vegetation drought response, we applied the extreme-based method (Sect. <xref ref-type="sec" rid="Ch1.S2.SS4"/>). This method identifies the dominant drought timescale, defined as the SPEI timescale (ranging from 1 to 24 months) that has the most negative impact on vegetation during drought periods.</p>

      <fig id="F5" specific-use="star"><label>Figure 5</label><caption><p id="d2e1710">Spatial patterns of the dominant drought timescale of GPP<sub>SA</sub> for <bold>(a)</bold> the MODIS reference and <bold>(b)</bold> the LPJML-5 model. The dominant timescale is defined as the shortest SPEI timescale (between 1 and 24 months) that shows the most negative GPP<sub>SA</sub> during drought periods (with a SPEI threshold of <inline-formula><mml:math id="M56" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>). Bare areas and sparse vegetation are filtered out. <bold>(c)</bold> Violin plot comparing the distribution of dominant drought timescales between MODIS and LPJmL-5.</p></caption>
          <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f05.png"/>

        </fig>

      <p id="d2e1757">The spatial distribution of dominant drought timescales, shown in Fig. <xref ref-type="fig" rid="F5"/>, shows a clear difference between MODIS (a) and the LPJmL-5 simulations (b). Compared to MODIS, LPJmL-5 exhibits shorter dominant timescales, indicating a faster GPP response to drought. This difference is most pronounced in the Southern Hemisphere. LPJmL-5 also shows a skewed distribution with a clear peak around 2 months (panel c), whereas the MODIS response is more evenly distributed. Overall, these patterns highlight differences in temporal sensitivity between LPJmL-5 and the satellite-derived reference, with LPJmL-5 showing shorter response times during drought periods.</p>
</sec>
<sec id="Ch1.S3.SS3">
  <label>3.3</label><title>MYD and ND response</title>
      <p id="d2e1770">After evaluating LPJmL-5's response to droughts in general, this study now shifts its focus to MYDs and NDs.</p>

      <fig id="F6" specific-use="star"><label>Figure 6</label><caption><p id="d2e1775">Timeseries of GPP<sub>SA</sub> for LPJmL-5 and MODIS. MYDs and NDs are shaded in red and orange, respectively. Correlation coefficients between LPJmL-5 and MODIS are shown for the full time series, MYDs, and NDs. The grey line indicates the SPEI-12. The red and blue dashed lines mark <inline-formula><mml:math id="M58" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M59" display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>, respectively.</p></caption>
          <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f06.png"/>

        </fig>


<sec id="Ch1.S3.SS3.SSS1">
  <label>3.3.1</label><title>Temporal patterns</title>
      <p id="d2e1823">Firstly, we examine how well LPJmL-5 captures the timing and magnitude of GPP<sub>SA</sub> during MYDs and NDs across the focus regions. Figure <xref ref-type="fig" rid="F6"/> shows the time series of GPP<sub>SA</sub> for the satellite-derived reference (MODIS) and LPJmL-5, with MYDs and NDs indicated by red and orange shaded areas, respectively.</p>
      <p id="d2e1846">Overall, LPJmL-5 captures the timing and magnitude of GPP<sub>SA</sub> anomalies reasonably well, showing relatively high correlations with MODIS in most regions. The model aligns particularly well in CAL, WEU, ARG, and AUS, while performance is somewhat lower in SA and BRA. Correlations during MYDs are higher than those for the entire time series in all regions except BRA, and generally higher than those during NDs, except in ARG and BRA. This suggests that LPJmL-5 simulates MYD dynamics more accurately than ND dynamics.</p>
      <p id="d2e1858">Nonetheless, some systematic differences remain during MYDs: (1) the model tends to simulate a too fast and severe decline in GPP<sub>SA</sub> at the onset of MYDs, and (2) it exhibits strong variability during MYD periods, especially in the form of abrupt recovery spikes that are not seen in the satellite-derived reference.</p>
      <p id="d2e1870">These biases are reflected across the focus regions to varying degrees. In WEU, the model simulates an early GPP<sub>SA</sub> drop that precedes the reference onset. CAL shows a similar offset, likely linked to the timing mismatch seen in Fig. <xref ref-type="fig" rid="F4"/>. In BRA, the first MYD minimum occurs too early in the model, while the second MYD ends with a sudden increase in GPP<sub>SA</sub> not seen in the reference, again indicating an early response and rapid recovery. ARG captures drought severity relatively well but still shows some abrupt recovery spikes during MYDs. SA also shows the rapid onset and overestimated recovery rate, despite persistently low SPEI-12 values. AUS shows the best overall agreement, with most MYDs being captured well. However, the second MYD still shows the strong variability mid-drought, though less severely.</p>
      <p id="d2e1894">Overall, while LPJmL-5 tends to simulate an overall more dynamic response with faster declines and rapid recovery during MYDs, it successfully reproduces the key temporal dynamics of MYDs relative to the satellite-derived reference.</p>

      <fig id="F7" specific-use="star"><label>Figure 7</label><caption><p id="d2e1899">Spatial pattern of the mean GPP<sub>SA</sub> during MYDs for <bold>(a)</bold> MODIS and <bold>(b)</bold> LPJmL-5. Positive values indicate higher GPP than normal during MYDs, while negative values indicate reduced GPP values. Bare areas and sparse vegetation are filtered out. <bold>(c)</bold> shows a violin plot comparing the distributions of the mean GPP<sub>SA</sub> during MYDs from MODIS and LPJmL-5.</p></caption>
            <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f07.png"/>

          </fig>

</sec>
<sec id="Ch1.S3.SS3.SSS2">
  <label>3.3.2</label><title>Spatial patterns</title>
      <p id="d2e1943">In addition to the regional time series analysis, we now examine global spatial patterns of vegetation response to MYDs. Figure <xref ref-type="fig" rid="F7"/> shows the spatial distribution of the mean GPP<sub>SA</sub> during MYDs, as derived from (a) the MODIS reference and (b) the LPJmL-5 model. Negative values (red) indicate below-average GPP during MYD periods, while positive values (green) indicate GPP being higher than average.</p>
      <p id="d2e1957">Overall, LPJmL-5 simulates less severe GPP<sub>SA</sub> during MYDs compared to MODIS. However, the Spearman rank correlation (0.326, <inline-formula><mml:math id="M70" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.001</mml:mn></mml:mrow></mml:math></inline-formula>) still indicates a similarity in the overall spatial patterns between the model and the reference. This means that the model captures some important hotspot areas, but there are still noticeable differences. We attribute these differences largely to the abrupt recovery events in the model, as seen in Fig. <xref ref-type="fig" rid="F6"/>. The violin plot (Fig. <xref ref-type="fig" rid="F7"/>c) shows that MODIS has a slightly lower median. However, LPJmL-5 displays more extreme values at both ends of the distribution, indicating greater variability in the modelled MYD responses.</p>

      <fig id="F8" specific-use="star"><label>Figure 8</label><caption><p id="d2e1987">Spatial differences in mean GPP<sub>SA</sub> during MYDs between MODIS and LPJmL-5. The map highlights regions where LPJmL-5 shows weaker or stronger vegetation responses relative to the satellite-derived reference. Purple shades indicate underestimated negative impacts, while orange shades indicate underestimated positive or overestimated negative impacts. White, grey and black shades represent regions where the model and reference agree closely on vegetation response during MYDs. Insets display two-dimensional kernel density estimates (KDEs) for seven vegetation types, illustrating the distribution of differences in mean MYD responses. KDE levels are set to [0.05, 0.15, 0.25, 0.5, 0.75, 0.9, 1.0]. Root mean square error (RMSE) for each plant functional type is included in the insets.</p></caption>
            <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f08.png"/>

          </fig>

      <p id="d2e2006">To further highlight these differences, Fig. <xref ref-type="fig" rid="F8"/> shows the spatial difference in mean GPP<sub>SA</sub> during MYDs between MODIS and LPJmL-5 and two-dimensional Kernel Density Estimate (KDE) plots for the different PFTs. Purple shades indicate underestimation of negative drought impacts, while orange shades indicate underestimation of positive impacts or overestimation of negative impacts. Black, grey and white areas indicate close agreement between the model and the reference.</p>
      <p id="d2e2020">LPJmL-5 shows a reasonable agreement with MODIS in most regions, as indicated by the black, grey and white areas. These areas include Central North America, Europe, and parts of East and West Asia. However, there are several regions that emerge as hotspots of LPJmL-5 underestimation of GPP anomalies (visible as dark purple areas), including Australia, Northwest India, Pakistan, Southern Africa, Central Argentina, Eastern Brazil, and Northern Mexico. In these regions, LPJmL-5 shows weaker or absent negative responses, suggesting that the model does not fully capture drought severity in these areas. This finding is consistent with the temporal patterns shown in Fig. <xref ref-type="fig" rid="F4"/>. Additionally, LPJmL-5 tends to underestimate the satellite-derived positive GPP responses in the arctic boreal zone (visible as orange areas). Here, the model does not reproduce the positive GPP anomalies during MYDs seen in the MODIS reference data (Fig. <xref ref-type="fig" rid="F7"/> and <xref ref-type="bibr" rid="bib1.bibx42" id="altparen.57"/>).</p>
      <p id="d2e2030">Beyond spatial patterns, differences between LPJmL-5 and MODIS also vary by PFT (Fig. <xref ref-type="fig" rid="F8"/>). Tropical trees show a wide spread in MYD GPP responses, while temperate trees align more closely with MODIS but still occasionally underestimate negative impacts (lower right quadrant). Grasslands show reasonable agreement, with LPJmL-5 capturing most directions of the MYD response correctly. Croplands, although displaying the weakest agreement in absolute GPP (Fig. <xref ref-type="fig" rid="F3"/>), show strong agreement in drought-induced GPP anomalies. This is reflected by the lowest RMSE among all PFTs (Fig. <xref ref-type="fig" rid="F8"/>). This indicates that, despite limitations in simulating absolute productivity, LPJmL-5 more consistently captures the relative response of croplands to drought events.</p>
      <p id="d2e2039">Overall, LPJmL-5 reproduces several key temporal and spatial MYD patterns present in the MODIS reference, but still shows some model bias in both time and space. Temporally, LPJmL-5 often simulates faster and stronger vegetation declines at the onset of the MYD than the satellite-derived reference and shows a fast recovery during the drought period. Spatially, LPJmL-5 shows a reasonable agreement with MODIS in most regions and for most PFTs, but also shows some underestimation of both strong negative and strong positive vegetation responses. These biases may be caused by short-term overreactions and rapid recoveries in the model's time series, which reduces the average drought response.</p>
</sec>
</sec>
</sec>
<sec id="Ch1.S4">
  <label>4</label><title>Discussion</title>
      <p id="d2e2052">Overall, this study assesses LPJmL-5's ability to simulate global vegetation responses to (multi-year) droughts. First, comparing dynamic and prescribed land cover settings in LPJmL-5 revealed that prescribed land cover improves agreement with MODIS GPP. The lowest agreement is generally found in the Southern Hemisphere and in cropland-dominated regions. Second, LPJmL-5 and MODIS GPP exhibit similar broad seasonal and interannual variability, although the model tends to simulate stronger GPP dynamics and introduces temporal shifts relative to the satellite-derived reference. Third, using the extreme-based method to identify the timescale at which drought has the strongest impact on vegetation, we found that LPJmL-5 simulates shorter dominant drought timescales than MODIS, particularly in tropical regions and the Southern Hemisphere, indicating a faster modelled GPP response to drought conditions. Fourth, the analysis of MYDs revealed a consistent pattern of model bias across both time and space. Temporally, LPJmL-5 simulates stronger and more rapid GPP declines at the onset of MYDs and shows faster recovery during drought periods compared to the reference. Spatially, the model shows muted representations of both strong negative and strong positive GPP anomalies compared to MODIS.</p>
      <p id="d2e2055">Overall, LPJmL-5 simulates faster GPP declines and recoveries during droughts than the satellite-derived reference, suggesting limitations in how the model represents vegetation resistance and resilience to water-induced stress and mortality.</p>
<sec id="Ch1.S4.SS1">
  <label>4.1</label><title>Uncertainties in the model choices and setup</title>
      <p id="d2e2065">Different choices in model parametrisation and setup can lead to significant differences in modelled GPP. For example, both the choice of meteorological forcing dataset (ERA5 <xref ref-type="bibr" rid="bib1.bibx19" id="paren.58"/> vs. W5E5 <xref ref-type="bibr" rid="bib1.bibx28" id="paren.59"/>) and the temporal resolution (monthly vs. daily) have a significant impact on simulated GPP patterns (Sect. S4). This indicates that it is important for future drought oriented studies to make clear choices with regard to the modelling setup. In this study, we selected the daily W5E5 dataset <xref ref-type="bibr" rid="bib1.bibx28" id="paren.60"/>, as it is bias-corrected and offers a high temporal resolution and the total global GPP was closest to that found in observational datasets.</p>
      <p id="d2e2077">Our analyses also showed that GPP output is highly sensitive to the radiation inputs (Sect. S4). GPP values varied substantially depending on whether the model calculated radiation internally or used observed radiation data. We found that errors in GPP are higher from internal estimation, compared to directly providing longwave and shortwave radiation from observations.</p>
      <p id="d2e2080">Using dynamic land cover in LPJmL-5 resulted in systematically overestimated GPP values across most regions (Fig. <xref ref-type="fig" rid="F2"/> and Sect. S5), also noted in <xref ref-type="bibr" rid="bib1.bibx46" id="text.61"/>. We argue that this difference is the result of a mismatch between modelled and observed vegetation distributions (Appendix C and Sect. S5), which is to be expected as the land cover is not constrained to the observed land cover. For example, LPJmL-5 simulates tree cover in parts of Western Europe where forests could exist naturally, but are absent today due to long-term human management. To better match satellite-derived GPP and reduce these discrepancies, we prescribed land cover fractions, rather than letting the model simulate land cover dynamically.</p>
</sec>
<sec id="Ch1.S4.SS2">
  <label>4.2</label><title>Evaluation of GPP simulations and vegetation drought response in LPJmL-5</title>
      <p id="d2e2096">Prescribing land cover in LPJmL-5 improved the agreement with MODIS GPP, as indicated by higher KGE values in Fig. <xref ref-type="fig" rid="F2"/>. This is consistent with studies showing that accurate land cover representation is essential for reproducing observed productivity patterns <xref ref-type="bibr" rid="bib1.bibx27" id="paren.62"/>, and that constraining DGVM simulations with satellite-derived observations (including land cover) enhances their ability to capture observed spatial patterns of vegetation and carbon dynamics <xref ref-type="bibr" rid="bib1.bibx14" id="paren.63"/>.</p>
      <p id="d2e2107">Breaking down the differences between LPJmL-5 and MODIS GPP during MYDs by vegetation type further highlights a varying model performance (Fig. <xref ref-type="fig" rid="F8"/>). Tropical trees show the largest variability in drought response, showing both under- and overestimations of impacts. Similar results were found by <xref ref-type="bibr" rid="bib1.bibx38" id="text.64"/>, who showed that terrestrial biosphere models underestimate drought impacts on tropical forest biomass. Temperate trees generally align more closely with observations but still underestimate negative drought impacts. This is consistent with <xref ref-type="bibr" rid="bib1.bibx26" id="text.65"/>, who found that land carbon models underestimate the severity and duration of drought effects on productivity across temperate forests in the US and Europe. Similarly, <xref ref-type="bibr" rid="bib1.bibx45" id="text.66"/> observed that models frequently overpredict GPP under dry conditions, further pointing to an underestimation of drought stress. Boreal trees and grasses are more prone to underestimated positive responses. Croplands, despite poorer agreement in absolute GPP dynamics, show comparatively better agreement in drought-induced anomalies. This suggests that while LPJmL-5 struggles to reproduce the magnitude of the GPP, it is able to capture relative deviations during drought events. Such behaviour indicates that biases in absolute GPP do not necessarily translate into errors in anomaly-based responses.</p>
      <p id="d2e2121">A key limitation underlying these issues appears to be how vegetation mortality is represented in LPJmL-5. In the model, mortality is only driven by low productivity and heat stress, and can not be directly induced by prolonged water deficits. Drought-induced mortality occurs only if plants fail to produce enough biomass to sustain themselves. However, this natural mortality is capped by a maximum annual rate <xref ref-type="bibr" rid="bib1.bibx47" id="paren.67"/>, which restricts how much vegetation can be lost due to die-off each year. Moreover, this mortality mechanism is only active in the dynamic land cover setting, and is not applied when land cover is prescribed. Since the simulations in this study use the prescribed land cover setting, no vegetation is lost due to this background mortality. This means that while GPP may decline sharply during droughts, vegetation does not die, causing unrealistically rapid recovery periods when precipitation (temporarily) returns. Although LPJmL-5 includes a heat-stress mortality mechanism, it only applies to boreal trees <xref ref-type="bibr" rid="bib1.bibx47" id="paren.68"/>. As a result, vegetation decline during MYDs is not directly driven by water shortages, which prevents large-scale die-off and causes the model to underestimate total vegetation loss. These limitations reduce the model's ability to simulate the full severity of vegetation decline, especially during MYDs. This issue has been noted by <xref ref-type="bibr" rid="bib1.bibx46" id="text.69"/>, who reported that the lack of realistic mortality processes leads to an overestimation of simulated biomass and contributes to biases in GPP. Similarly, <xref ref-type="bibr" rid="bib1.bibx31" id="text.70"/> found that models which combine carbon starvation with hydraulic failure mechanisms are better able to reproduce observed drought-induced tree mortality, and <xref ref-type="bibr" rid="bib1.bibx67" id="text.71"/> highlighted the need to account for both stomatal and non-stomatal limitations to realistically capture plant responses to water stress. <xref ref-type="bibr" rid="bib1.bibx33" id="text.72"/> further showed that including hydraulic processes in LPJ-GUESS improves simulations of drought responses. Together, these findings suggest that LPJmL-5 likely underestimates the true impact of drought on vegetation.</p>
      <p id="d2e2143">Despite underestimating long-term vegetation loss, LPJmL-5 tends to overestimate vegetation decline at the start of droughts, as shown in Figs. <xref ref-type="fig" rid="F5"/> and <xref ref-type="fig" rid="F6"/>. LPJmL-5 does include a water stress factor that influences the growth, which results in a rapid decline in GPP. This stress is determined by soil moisture availability and fixed rooting depths for each PFT, without accounting for dynamic rooting strategies, capillary rise or explicit groundwater processes <xref ref-type="bibr" rid="bib1.bibx47" id="paren.73"/>. However, in reality, vegetation often withstands short-term stress periods through drought survival strategies <xref ref-type="bibr" rid="bib1.bibx44 bib1.bibx37" id="paren.74"/>. At the same time, LPJmL-5 also simulates unrealistic recoveries during the drought period. Since there is no mortality, vegetation can survive during a drought period, resulting in no loss of biomass. When conditions improve temporarily, for example due to brief rainfall, the model quickly simulates regrowth. This leads to an unrealistic drought recovery effect. This is in contrast with real vegetation, which often suffers from long-term impacts <xref ref-type="bibr" rid="bib1.bibx62" id="paren.75"/>. Both the quick decline and recovery are consistent with findings by <xref ref-type="bibr" rid="bib1.bibx26" id="text.76"/>, who show that land carbon models underestimate the severity and duration of drought impact on plant productivity.</p>
      <p id="d2e2164">Together, the combination of overly sensitive short-term GPP decline, limited representation of long-term vegetation loss, and unrealistically fast recovery during drought periods results in a mismatch in LPJmL-5's simulated vegetation drought response. While prescribing land cover improves the overall GPP performance, these structural issues in the mortality representation limit the model's ability to simulate realistic vegetation dynamics during drought events. This is especially limiting when the analysis is focused on multi-year drought events, which have been shown to cause higher vegetation mortality rates <xref ref-type="bibr" rid="bib1.bibx5" id="paren.77"/> and are expected to become more frequent under climate change <xref ref-type="bibr" rid="bib1.bibx59" id="paren.78"/>.</p>
</sec>
<sec id="Ch1.S4.SS3">
  <label>4.3</label><title>Uncertainties in methodological choices</title>
      <p id="d2e2182">To improve the overall quality and reliability of our simulations, we used the state-of-the-art dynamic vegetation model LPJmL-5. This model is publicly available with an extensive model description and evaluation <xref ref-type="bibr" rid="bib1.bibx47 bib1.bibx46 bib1.bibx56" id="paren.79"/>. LPJmL-5 provides a multitude of vegetation-related outputs on ecological, hydrological, and agricultural components, making it suitable for studying vegetation responses at the global scale.</p>
      <p id="d2e2188">Drought events were identified using the Standardized Precipitation Evapotranspiration Index (SPEI), which incorporates both precipitation (<inline-formula><mml:math id="M73" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula>) and potential evapotranspiration (PET), allowing it to better reflect the impact of temperature on drought severity <xref ref-type="bibr" rid="bib1.bibx54" id="paren.80"/>. As a standardized index, SPEI also enables consistent spatial comparisons of drought conditions at the global scale. By using a 12 month aggregation period for the SPEI, we also remove seasonal variations and focus on capturing MYDs.</p>
      <p id="d2e2201">As a standardized metric, SPEI characterizes drought conditions relative to the local climatic baseline, reflecting periods that are characterized by below normal water availability. In energy-limited regions, such as the tropics or boreal forests, negative SPEI values may occur even when absolute water availability remains relatively high <xref ref-type="bibr" rid="bib1.bibx66" id="paren.81"/>. To ensure these droughts correspond to meaningful reductions in water availability, we compared soil water content (SWC) during droughts and normal conditions (Sect. S2), finding consistently lower SWC across soil layers. This supports the interpretation that identified MYDs correspond to periods of meaningful vegetation stress, even in energy-limited regions.</p>
      <p id="d2e2207">While there is currently no universally agreed-upon definition of MYDs, we follow the approach of <xref ref-type="bibr" rid="bib1.bibx59" id="text.82"/> (see Sect. <xref ref-type="sec" rid="Ch1.S2.SS1"/>), which was also adopted in <xref ref-type="bibr" rid="bib1.bibx42" id="text.83"/> and <xref ref-type="bibr" rid="bib1.bibx53" id="text.84"/>. SPEI values are sensitive to the method and input data used for calculating PET. We compared results using ERA5 and W5E5 and found differences in the number of MYDs identified for the period 2000–2019 (Sect. S4). Nonetheless, the MYDs identified in our focus regions are broadly consistent with those reported in other studies <xref ref-type="bibr" rid="bib1.bibx53 bib1.bibx59 bib1.bibx30 bib1.bibx29 bib1.bibx6 bib1.bibx36 bib1.bibx51" id="paren.85"/>, despite minor differences in timing.</p>
      <p id="d2e2225">To evaluate vegetation responses, we used monthly MODIS GPP as a proxy for ecosystem productivity, representing the amount of carbon captured from the atmosphere through photosynthesis <xref ref-type="bibr" rid="bib1.bibx3" id="paren.86"/>. Previous drought studies have often relied on the EVI <xref ref-type="bibr" rid="bib1.bibx42 bib1.bibx20 bib1.bibx64" id="paren.87"/>, but EVI is satellite-derived and not directly available from LPJmL-5. Because MODIS GPP and EVI are strongly correlated (<xref ref-type="bibr" rid="bib1.bibx39 bib1.bibx48" id="altparen.88"/>; see also Sect. S3), including during multi-year droughts <xref ref-type="bibr" rid="bib1.bibx42" id="paren.89"/>, MODIS GPP can be considered a suitable proxy for vegetation activity.</p>
      <p id="d2e2240">Several studies further indicate that MODIS GPP provides a meaningful representation of productivity across biomes, with most errors linked to input datasets rather than the algorithm itself. Performance is generally good in temperate and mixed forests, while errors are larger in evergreen broadleaf forests and tropical regions <xref ref-type="bibr" rid="bib1.bibx49" id="paren.90"/>. <xref ref-type="bibr" rid="bib1.bibx57" id="text.91"/> showed that much of the error  originates from FPAR, land-cover misclassification, and light-use efficiency parameters rather than meteorology. Systematic underestimation has also been reported in grasslands <xref ref-type="bibr" rid="bib1.bibx68" id="paren.92"/>. These studies agree that cloud contamination and uncertainties in tropical regions remain key challenges. Taken together, the strong correspondence between MODIS GPP, EVI, and solar-induced fluorescence (Sect. S3) supports the use of MODIS GPP as our validation dataset for LPJmL-5 GPP, while acknowledging that performance is weaker in tropical ecosystems and that results should be interpreted with these limitations in mind.</p>
      <p id="d2e2252">Although other sources of GPP data exist, such as site-level flux tower measurements, we choose to use satellite-derived MODIS GPP because it provides consistent, observation-based estimates at the global scale. To maintain observational consistency and avoid mixing in other models, we did not include GPP estimates derived from other land surface models as they are likely based on similar assumptions and equations as the LPJmL-5 model. Nonetheless, MODIS GPP is subject to uncertainty, particularly in tropical regions (Sect. S3), and this should be kept in mind when interpreting the results.</p>
</sec>
</sec>
<sec id="Ch1.S5" sec-type="conclusions">
  <label>5</label><title>Conclusion</title>
      <p id="d2e2265">This study evaluated LPJmL-5's ability to simulate global vegetation responses to drought using MODIS-derived GPP as a reference. Our analysis shows that LPJmL-5 simulates shorter response times to drought than observed by MODIS, particularly in tropical regions and the Southern Hemisphere. During MYDs, LPJmL-5 captures the key temporal and spatial dynamics found in MODIS. However, the model tends to overestimate vegetation response at the MYD onset and shows rapid recovery behaviour. This results in muted overall drought impacts and an underrepresentation of both strong negative and strong positive GPP anomalies.</p>
      <p id="d2e2268">Among PFTs, croplands show the best agreement with satellite-derived MYD responses. In contrast, boreal vegetation shows underestimated positive drought responses and temperate vegetation underestimated negative ones. Tropical vegetation displays more mixed results, with both over- and underestimation. These differences between modelled and satellite-derived GPP during drought periods may be partly attributed to how LPJmL-5 represents vegetation stress and drought-induced mortality. In particular, while the model responds strongly to short-term reductions in productivity, it limits long-term vegetation decline, leading to rapid recovery and reduced vegetation die-off during drought periods.</p>
      <p id="d2e2271">Beyond drought conditions, prescribing land cover instead of simulating it dynamically improves the model's ability to reproduce global GPP dynamics. Overall, LPJmL-5 reproduces GPP reasonably well, although performance declines in parts of the Southern Hemisphere and in cropland-dominated regions. This suggests that general GPP simulation performance is not necessarily linked to performance during drought conditions.</p>
      <p id="d2e2274">In conclusion, LPJmL-5 is able to reproduce several key features of vegetation responses to MYDs, but tends to simulate vegetation that is too responsive during both the drought onset and the recovery. This points to limitations in the representation of vegetation resistance and resilience to water stress and mortality. Improving these processes is important to more accurately capture the magnitude of drought impacts and for strengthening confidence in model-based assessments of vegetation responses to droughts.</p>
</sec>

      
      </body>
    <back><app-group>

<app id="App1.Ch1.S1">
  <label>Appendix A</label><title>Dominant land cover types</title>
      <p id="d2e2289">Figure <xref ref-type="fig" rid="FA1"/> shows the dominant land cover types at a 0.1° spatial resolution, based on LPJmL-5's land cover and land use dataset (see Table <xref ref-type="table" rid="T1"/>). Cropland fractions were combined into a single “cropland” category, while natural PFTs were grouped into broader vegetation classes: tropical trees (tropical broadleaved evergreen and raingreen trees), temperate trees (needleleaved evergreen, broadleaved evergreen, and broadleaved summergreen trees), boreal trees (needleleaved evergreen, broadleaved summergreen, and needleleaved summergreen trees), tropical C<sub>4</sub> grass, temperate C<sub>3</sub> grass, and polar C<sub>3</sub> grass.</p>

      <fig id="FA1"><label>Figure A1</label><caption><p id="d2e2325">Dominant land cover types at a 0.1° resolution for 2019. Vegetation categories include tropical trees (TTr), temperate trees (TTe), boreal trees (TBo), cropland (Cr), tropical C<sub>4</sub> grass (GTr), temperate C<sub>3</sub> grass (GTe), and polar C<sub>3</sub> grass (GBo).</p></caption>
        
        <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f09.png"/>

      </fig>


</app>

<app id="App1.Ch1.S2">
  <label>Appendix B</label><title>Number of MYDs and chosen focus regions</title>
      <p id="d2e2373">Figure <xref ref-type="fig" rid="FB1"/> shows the number of MYDs (2000–2019) at a 0.5° spatial resolution from W5E5 data <xref ref-type="bibr" rid="bib1.bibx28" id="paren.93"/> (see Sects. 2.1 and S1). Black outlines indicate the chosen focus regions: California (CAL), the Rhine-Meuse delta in western Europe (WEU), the Brahmaputra River basin in Bangladesh/India/Bhutan/China (BRA), central Argentina (ARG), the Orange River basin in southern Africa (SA), and the Murray-Darling basin in Australia (AUS) (see Sect. 2.6).</p>

      <fig id="FB1"><label>Figure B1</label><caption><p id="d2e2383">Number of MYDs between 2000–2019 at 0.5° spatial resolution. Bare areas and regions with sparse vegetation are excluded. Black contours indicate the six chosen focus regions: CAL, WEU, BRA, ARG, SA, and AUS.</p></caption>
        
        <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f10.png"/>

      </fig>


</app>

<app id="App1.Ch1.S3">
  <label>Appendix C</label><title>LPJmL dynamic vs. ESA prescribed land cover</title>
      <p id="d2e2405">Figure <xref ref-type="fig" rid="FC1"/> shows the difference in vegetation cover fractions between LPJmL-5 and ESA. Globally, the ESA dataset shows greater coverage of temperate and polar C<sub>3</sub> grasses, as well as boreal needleleaved summergreen forests. In contrast, LPJmL-5's dynamic vegetation simulates more temperate needleleaved evergreen trees, boreal needleleaved evergreen trees, boreal broadleaved summergreen trees, and tropical C<sub>4</sub> grasses.</p>
      <p id="d2e2428">The CDFs in Fig. <xref ref-type="fig" rid="FC2"/> further show the impact of land cover on model performance. Across all vegetation types, the prescribed land cover shows higher KGE values. The difference between prescibed and dynamic land cover is particularly pronounced for temperate C<sub>3</sub> grasses, which perform poorly with the the dynamic land cover. LPJmL-5 substantially underrepresents the spatial extent of temperate C<sub>3</sub> grasses, assigning them to only 2.7 % of grid cells, compared to 22.2 % under the ESA-based classification. This mismatch likely contributes to the poor performance, as misclassified vegetation types lead to unrealistic GPP estimates. Tree-dominated grid cells also show better performance with prescribed land cover, however, the difference is less severe than for grasses.</p><fig id="FC1"><label>Figure C1</label><caption><p id="d2e2453">Difference in vegetation cover fractions between LPJmL-5 and ESA. Blue areas indicate where ESA has a higher vegetation fraction; red areas indicate where LPJmL-5 simulates a higher vegetation fraction.</p></caption>
        
        <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f11.png"/>

      </fig>

<fig id="FC2"><label>Figure C2</label><caption><p id="d2e2468">Cumulative distribution function (CDF) of the KGE of the <bold>(a)</bold> prescribed land cover GPP and <bold>(b)</bold> dynamic land cover for the different vegetation types in LPJmL-5. Percentages show the percentage of grid cells with that land cover. Black dashed line indicates KGE <inline-formula><mml:math id="M84" display="inline"><mml:mrow><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.41</mml:mn></mml:mrow></mml:math></inline-formula>. Values greater than <inline-formula><mml:math id="M85" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.41</mml:mn></mml:mrow></mml:math></inline-formula> indicate that the model improved upon the mean benchmark.</p></caption>
        
        <graphic xlink:href="https://esd.copernicus.org/articles/17/607/2026/esd-17-607-2026-f12.png"/>

      </fig>

</app>
  </app-group><notes notes-type="codedataavailability"><title>Code and data availability</title>

      <p id="d2e2511">The LPJmL-5 source code is openly available at <uri>https://github.com/PIK-LPJmL/LPJmL</uri> (last access: 7 April 2025) and <ext-link xlink:href="https://doi.org/10.5281/zenodo.14012503" ext-link-type="DOI">10.5281/zenodo.14012503</ext-link> <xref ref-type="bibr" rid="bib1.bibx60" id="paren.94"/>. This study uses version v5.7.9 (released on 2 May 2024).</p>

      <p id="d2e2523">All processed data required to reproduce the analyses are available at: <ext-link xlink:href="https://doi.org/10.24416/UU01-U97GDE" ext-link-type="DOI">10.24416/UU01-U97GDE</ext-link> <xref ref-type="bibr" rid="bib1.bibx41" id="paren.95"/>.</p>

      <p id="d2e2532">The raw datasets are publicly available from their respective sources (see Table <xref ref-type="table" rid="T1"/>).</p>

      <p id="d2e2537">All scripts used for postprocessing and analysis in this study are available at <ext-link xlink:href="https://doi.org/10.5281/zenodo.17085725" ext-link-type="DOI">10.5281/zenodo.17085725</ext-link> <xref ref-type="bibr" rid="bib1.bibx40" id="paren.96"/>.</p>
  </notes><app-group>
        <supplementary-material position="anchor"><p id="d2e2546">The supplement related to this article is available online at <inline-supplementary-material xlink:href="https://doi.org/10.5194/esd-17-607-2026-supplement" xlink:title="pdf">https://doi.org/10.5194/esd-17-607-2026-supplement</inline-supplementary-material>.</p></supplementary-material>
        </app-group><notes notes-type="authorcontribution"><title>Author contributions</title>

      <p id="d2e2555">DR: Conceptualisation, methodology, software, validation, formal analysis, data curation, writing – original draft, visualisation. SH: Conceptualisation, formal analysis, writing – review and editing. HB: Conceptualisation, formal analysis, writing – review and editing. NW: Conceptualisation, formal analysis, writing – review and editing, project administration, funding acquisition.</p>
  </notes><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d2e2561">The contact author has declared that none of the authors has any competing interests.</p>
  </notes><notes notes-type="disclaimer"><title>Disclaimer</title>

      <p id="d2e2569">Views and opinions expressed are those of the author(s) only and do not necessarily reflect those of the European Union or the European Research Council. Neither the European Union nor the granting authority can be held responsible for them.Publisher's note: Copernicus Publications remains neutral with regard to jurisdictional claims made in the text, published maps, institutional affiliations, or any other geographical representation in this paper. The authors bear the ultimate responsibility for providing appropriate place names. Views expressed in the text are those of the authors and do not necessarily reflect the views of the publisher.</p>
  </notes><notes notes-type="financialsupport"><title>Financial support</title>

      <p id="d2e2578">This research has been supported by the European Research Council, H2020 European Research Council (grant no. 101075354). DR and NW are supported by the ERC STG MultiDry project. SH is supported by NWO VIDI project Recover VI.Vidi.223.009. HB is supported by the European Union (ERC-CoG, 3POLE2SEA, 101126168).</p>
  </notes><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d2e2585">This paper was edited by Lan Wang-Erlandsson and reviewed by Xiangyi Li and one anonymous referee.</p>
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