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  <front>
    <journal-meta><journal-id journal-id-type="publisher">ESD</journal-id><journal-title-group>
    <journal-title>Earth System Dynamics</journal-title>
    <abbrev-journal-title abbrev-type="publisher">ESD</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Earth Syst. Dynam.</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">2190-4987</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/esd-16-1569-2025</article-id><title-group><article-title>100 kyr ice age cycles as a timescale-matching problem</article-title><alt-title>100 <inline-formula><mml:math id="M1" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> ice age cycles as a timescale-matching problem</alt-title>
      </title-group>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1">
          <name><surname>Mitsui</surname><given-names>Takahito</given-names></name>
          <email>takahito321@gmail.com</email>
        <ext-link>https://orcid.org/0000-0002-2825-3996</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>Ditlevsen</surname><given-names>Peter</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-2120-7732</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff3 aff4">
          <name><surname>Boers</surname><given-names>Niklas</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff5">
          <name><surname>Crucifix</surname><given-names>Michel</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-3437-4911</ext-link></contrib>
        <aff id="aff1"><label>1</label><institution>Faculty of Health Data Science, Juntendo Univerity, Urayasu, Chiba, Japan</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Niels Bohr Institute, University of Copenhagen, Copenhagen, Denmark</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Earth System Modelling, School of Engineering and Design, Technical University of Munich, Munich, Germany</institution>
        </aff>
        <aff id="aff4"><label>4</label><institution>Potsdam Institute for Climate Impact Research, Member of the Leibniz Association, Potsdam, Germany</institution>
        </aff>
        <aff id="aff5"><label>5</label><institution>Earth and Life Institute, UCLouvain, Louvain-la-Neuve, Belgium</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Takahito Mitsui (takahito321@gmail.com)</corresp></author-notes><pub-date><day>26</day><month>September</month><year>2025</year></pub-date>
      
      <volume>16</volume>
      <issue>5</issue>
      <fpage>1569</fpage><lpage>1584</lpage>
      <history>
        <date date-type="received"><day>2</day><month>December</month><year>2024</year></date>
           <date date-type="accepted"><day>18</day><month>July</month><year>2025</year></date>
           <date date-type="rev-recd"><day>8</day><month>July</month><year>2025</year></date>
           <date date-type="rev-request"><day>6</day><month>December</month><year>2024</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2025 Takahito Mitsui et al.</copyright-statement>
        <copyright-year>2025</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025.html">This article is available from https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025.html</self-uri><self-uri xlink:href="https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025.pdf">The full text article is available as a PDF file from https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025.pdf</self-uri>
      <abstract><title>Abstract</title>

      <p id="d2e146">The dominant period of the Late Pleistocene glacial–interglacial cycles is roughly 100 <inline-formula><mml:math id="M2" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, rather than other major astronomical periods such as 19, 23, 41, and 400 <inline-formula><mml:math id="M3" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. Various models explain this fact through distinct dynamical mechanisms, including synchronization of self-sustained oscillations and resonance in mono- or multi-stable systems. However, the diversity of proposed models and dynamical mechanisms may obscure the essential factor behind the emergence of the <inline-formula><mml:math id="M4" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M5" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity. We propose the hypothesis that the ice-sheet climate system responds to astronomical forcing at the <inline-formula><mml:math id="M6" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M7" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity because the intrinsic timescale of the system is closer to 100 <inline-formula><mml:math id="M8" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> than to other major astronomical periods. We support this idea with analyses and sensitivity studies of several simple ice age models with contrasting mechanisms.</p>
  </abstract>
    
<funding-group>
<award-group id="gs1">
<funding-source>Volkswagen Foundation</funding-source>
<award-id>na</award-id>
</award-group>
<award-group id="gs2">
<funding-source>H2020 Marie Skłodowska-Curie Actions</funding-source>
<award-id>956170</award-id>
</award-group>
<award-group id="gs3">
<funding-source>Bundesministerium für Bildung und Forschung</funding-source>
<award-id>01LS3001A</award-id>
</award-group>
<award-group id="gs4">
<funding-source>Japan Society for the Promotion of Science</funding-source>
<award-id>25K07942</award-id>
</award-group>
<award-group id="gs5">
<funding-source>HORIZON EUROPE Framework Programme</funding-source>
<award-id>101137601</award-id>
</award-group>
</funding-group>
</article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction</title>
      <p id="d2e219">Glacial–interglacial cycles are a pronounced mode of climate variability in the Pleistocene, accompanied by large changes in temperatures <xref ref-type="bibr" rid="bib1.bibx16 bib1.bibx38" id="paren.1"/>, global ice volume <xref ref-type="bibr" rid="bib1.bibx71" id="paren.2"/>, and greenhouse gas concentrations <xref ref-type="bibr" rid="bib1.bibx7 bib1.bibx45" id="paren.3"/>. Changes in global ice volume are recorded, e.g., in the oxygen isotope ratio <inline-formula><mml:math id="M9" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> of benthic foraminifera found in marine sediments <xref ref-type="bibr" rid="bib1.bibx44" id="paren.4"/> (Fig. <xref ref-type="fig" rid="F1"/>d), where higher <inline-formula><mml:math id="M10" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> values indicate larger ice volume and lower deep-ocean temperatures. The dominant period of the Late Pleistocene glacial cycles is roughly 100 <inline-formula><mml:math id="M11" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, as shown in the power spectral density (PSD) (Fig. <xref ref-type="fig" rid="F1"/>f; see Appendix A for the PSD method).</p>

      <fig id="F1" specific-use="star"><label>Figure 1</label><caption><p id="d2e275">Time series and power spectral densities (PSDs) of the astronomical forcing <xref ref-type="bibr" rid="bib1.bibx40" id="paren.5"/> and glacial cycles over the last 1 <inline-formula><mml:math id="M12" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula>. <bold>(a)</bold> Obliquity. <bold>(b)</bold> Climatic precession (green) and eccentricity (magenta). <bold>(c)</bold> Summer solstice insolation at 65° N. <bold>(d)</bold> Benthic <inline-formula><mml:math id="M13" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> stack records representing glacial–interglacial cycles. The so-called LR04 record with orbital tuning (black) <xref ref-type="bibr" rid="bib1.bibx44" id="paren.6"/>, the LR04 record without orbital tuning (red) <xref ref-type="bibr" rid="bib1.bibx43" id="paren.7"/>, and the record without orbital tuning (blue) <xref ref-type="bibr" rid="bib1.bibx31" id="paren.8"/>. Note that the vertical axis is reversed so that larger <inline-formula><mml:math id="M14" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> values, corresponding to colder conditions, are lower. <bold>(e)</bold> PSD of the eccentricity (magenta) and the PSD of the summer solstice insolation <inline-formula><mml:math id="M15" display="inline"><mml:mrow><mml:msub><mml:mi>F</mml:mi><mml:mtext>65N</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (orange). <bold>(f)</bold> PSDs from each benthic <inline-formula><mml:math id="M16" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> record in <bold>(d)</bold>. The dashed vertical lines in <bold>(e)</bold> and <bold>(f)</bold> indicate major astronomical periods <xref ref-type="bibr" rid="bib1.bibx40" id="paren.9"/>.</p></caption>
        <graphic xlink:href="https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025-f01.png"/>

      </fig>

      <p id="d2e387">Summer insolation in the high northern latitudes (Fig. <xref ref-type="fig" rid="F1"/>c) is supposed to be a major driver <xref ref-type="bibr" rid="bib1.bibx49 bib1.bibx70" id="paren.10"/> or a pacemaker <xref ref-type="bibr" rid="bib1.bibx29" id="paren.11"/> of the glacial–interglacial cycles. It fluctuates due to long-term variations in the astronomical parameters: climatic precession <inline-formula><mml:math id="M17" display="inline"><mml:mrow><mml:mi>e</mml:mi><mml:mi>sin⁡</mml:mi><mml:mi mathvariant="italic">ϖ</mml:mi></mml:mrow></mml:math></inline-formula> (and co-precession <inline-formula><mml:math id="M18" display="inline"><mml:mrow><mml:mi>e</mml:mi><mml:mi>cos⁡</mml:mi><mml:mi mathvariant="italic">ϖ</mml:mi></mml:mrow></mml:math></inline-formula>) with 19, 22.4, and 23.7 <inline-formula><mml:math id="M19" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> dominant periods (Fig. <xref ref-type="fig" rid="F1"/>b, green); obliquity <inline-formula><mml:math id="M20" display="inline"><mml:mi mathvariant="italic">ε</mml:mi></mml:math></inline-formula> with a 41 <inline-formula><mml:math id="M21" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> period (Fig. <xref ref-type="fig" rid="F1"/>a); and eccentricity <inline-formula><mml:math id="M22" display="inline"><mml:mi>e</mml:mi></mml:math></inline-formula> with 95, 124, and 405 <inline-formula><mml:math id="M23" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periods (Fig. <xref ref-type="fig" rid="F1"/>b, magenta) <xref ref-type="bibr" rid="bib1.bibx40 bib1.bibx9" id="paren.12"/>, where the periods of eccentricity are linked with those of climatic precession by relations of combination tones such as <inline-formula><mml:math id="M24" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">95</mml:mn><mml:mo>≃</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">19</mml:mn><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">23.7</mml:mn></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx10" id="paren.13"/>. These astronomical periods are in fact imprinted in the PSDs of the <inline-formula><mml:math id="M25" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> records, as shown in Fig. <xref ref-type="fig" rid="F1"/>f <xref ref-type="bibr" rid="bib1.bibx29" id="paren.14"/>.</p>
      <p id="d2e522">However, despite the dominant average period of the Late Pleistocene glacial cycles being <inline-formula><mml:math id="M26" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M27" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, the boreal summer insolation only has negligible power at this frequency. This discrepancy is known as the 100 <inline-formula><mml:math id="M28" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> problem. Instead, boreal summer insolation exhibits strong power in the 19–23.7 <inline-formula><mml:math id="M29" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> precession band and the 41 <inline-formula><mml:math id="M30" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> obliquity band (Fig. <xref ref-type="fig" rid="F1"/>e, orange). Hence, the <inline-formula><mml:math id="M31" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M32" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> glacial cycles have previously been explained as a response to four or five precession cycles <xref ref-type="bibr" rid="bib1.bibx68 bib1.bibx14 bib1.bibx30" id="paren.15"/>, a response to two or three obliquity cycles <xref ref-type="bibr" rid="bib1.bibx34" id="paren.16"/>, or a combination thereof <xref ref-type="bibr" rid="bib1.bibx32 bib1.bibx76 bib1.bibx72" id="paren.17"/>. Note that a single response to four or five precession cycles generally coincides with a one-to-one response to <inline-formula><mml:math id="M33" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M34" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity cycles, since a deglaciation in response to climatic precession <inline-formula><mml:math id="M35" display="inline"><mml:mrow><mml:mi>e</mml:mi><mml:mi>sin⁡</mml:mi><mml:mi mathvariant="italic">ϖ</mml:mi></mml:mrow></mml:math></inline-formula> tends to occur near the rising limb of eccentricity <inline-formula><mml:math id="M36" display="inline"><mml:mi>e</mml:mi></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx66 bib1.bibx1" id="paren.18"/>. Thus, eccentricity seems to impact  the pace of glacial cycles via the modulation of climatic precession, even though the boreal summer insolation forcing only has negligible 100 <inline-formula><mml:math id="M37" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> power.</p>
      <p id="d2e646">Until now, we have referred to the dominant period of the Late Pleistocene glacial cycles as <inline-formula><mml:math id="M38" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M39" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. Examining the PSDs of the benthic <inline-formula><mml:math id="M40" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> stack records over the last 1 <inline-formula><mml:math id="M41" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx44" id="paren.19"/>, the <inline-formula><mml:math id="M42" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M43" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> spectral peak actually aligns with the 95 <inline-formula><mml:math id="M44" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity peak, and it is indeed distinct from other potential eccentricity peaks such as 124 <inline-formula><mml:math id="M45" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> (Fig. <xref ref-type="fig" rid="F1"/>e and f). Concerns could be raised about using a tuned record for such an assessment, but the same conclusion is also drawn from two other records that are free from orbital tuning (<xref ref-type="bibr" rid="bib1.bibx43" id="altparen.20"/>; <xref ref-type="bibr" rid="bib1.bibx31" id="altparen.21"/>). Thus, in this study, the 95 <inline-formula><mml:math id="M46" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> period is assumed as the strongest mode over the last 1 <inline-formula><mml:math id="M47" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula> (<xref ref-type="bibr" rid="bib1.bibx16" id="altparen.22"/>, and <xref ref-type="bibr" rid="bib1.bibx67" id="altparen.23"/>, also specifically state the 95 <inline-formula><mml:math id="M48" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> period). This strong imprint of the 95 <inline-formula><mml:math id="M49" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity period (i.e., the combination tone of the climatic precession periods 23.7 and 19 <inline-formula><mml:math id="M50" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>) is consistent with recent studies suggesting that the timings of deglaciations are more tightly coupled with multiple climatic precession cycles than multiple obliquity cycles with 82 or 123 <inline-formula><mml:math id="M51" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx30 bib1.bibx15 bib1.bibx1" id="paren.24"/>.</p>
      <p id="d2e793">Synchronization and nonlinear resonance are two major dynamical mechanisms that result in a system's response tightly coupled with external forcing. Due to their ubiquity in nature, they are often invoked to explain the emergence of <inline-formula><mml:math id="M52" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M53" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles. As they are central to the discussion that follows, we briefly review them below.</p>
      <p id="d2e814">In <italic>synchronization</italic> (a.k.a. <italic>frequency entrainment</italic>, <italic>phase locking</italic>, or <italic>frequency locking</italic>)<fn id="Ch1.Footn1"><p id="d2e829">In this study, we follow the definition of synchronization from <xref ref-type="bibr" rid="bib1.bibx63" id="text.25"/>, where the terms frequency entrainment, phase locking, and frequency locking are considered synonymous with synchronization, assuming the prior existence of an underlying self-sustained oscillation that is being “locked”:  <xref ref-type="bibr" rid="bib1.bibx63" id="text.26"/> explicitly distinguish these notions from resonance or nonlinear response. In many studies of glacial cycles, however, the term “phase locking” is used to describe both synchronization and the nonlinear response, regardless of the existence of underlying self-sustained oscillations.</p></fn>, the system is assumed to exhibit self-sustained oscillations in the absence of forcing, and the frequency of the underlying oscillations is adjusted to match one of the  frequencies of external forcing, its harmonics, subharmonics, or a combination of these <xref ref-type="bibr" rid="bib1.bibx63" id="paren.27"/>. Many ice age models generate <inline-formula><mml:math id="M54" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M55" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles through the synchronization mechanism <xref ref-type="bibr" rid="bib1.bibx74 bib1.bibx27 bib1.bibx2 bib1.bibx21 bib1.bibx18 bib1.bibx3 bib1.bibx56 bib1.bibx53 bib1.bibx54 bib1.bibx39" id="paren.28"/>. Synchronization occurs more easily when the frequency of external forcing is closer to the natural frequency of the system's underlying self-sustained oscillations <xref ref-type="bibr" rid="bib1.bibx63" id="paren.29"/>. Thus if the <inline-formula><mml:math id="M56" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M57" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles are realized via synchronization, it suggests the existence of underlying self-sustained oscillations at <inline-formula><mml:math id="M58" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M59" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> timescale.</p>
      <p id="d2e904"><italic>Resonance</italic>, on the other hand, refers to an enhanced output response to an external forcing. Linear resonance is well known, in which the response to a periodic forcing is amplified when the external frequency <inline-formula><mml:math id="M60" display="inline"><mml:mi mathvariant="normal">Ω</mml:mi></mml:math></inline-formula> matches the system's natural frequency <inline-formula><mml:math id="M61" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ω</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>. In nonlinear systems, more complex behavior called <italic>nonlinear resonances</italic> occurs, such as the following: (i) the resonant frequency <inline-formula><mml:math id="M62" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="normal">Ω</mml:mi><mml:mi mathvariant="normal">r</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> that yields the maximum response can deviate from the natural frequency <inline-formula><mml:math id="M63" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ω</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> as the forcing amplitude increases; (ii) resonances can occur at superharmonics <inline-formula><mml:math id="M64" display="inline"><mml:mrow><mml:mi>m</mml:mi><mml:mi mathvariant="normal">Ω</mml:mi></mml:mrow></mml:math></inline-formula>, subharmonics <inline-formula><mml:math id="M65" display="inline"><mml:mrow><mml:mi mathvariant="normal">Ω</mml:mi><mml:mo>/</mml:mo><mml:mi>n</mml:mi></mml:mrow></mml:math></inline-formula>, and supersubharmonics <inline-formula><mml:math id="M66" display="inline"><mml:mrow><mml:mi>m</mml:mi><mml:mi mathvariant="normal">Ω</mml:mi><mml:mo>/</mml:mo><mml:mi>n</mml:mi></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M67" display="inline"><mml:mrow><mml:mi>m</mml:mi><mml:mo>,</mml:mo><mml:mi>n</mml:mi><mml:mo>∈</mml:mo><mml:mi mathvariant="double-struck">N</mml:mi></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx37" id="paren.30"/>. Recently, the term <italic>resonance</italic> has been generalized to include a broader range of processes that involve the enhancement, suppression, or optimization of a system's response through the variation, perturbation, or modulation of any system property <xref ref-type="bibr" rid="bib1.bibx65" id="paren.31"/>. In nonlinear resonance mechanisms of <inline-formula><mml:math id="M68" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M69" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> ice age cycles, the underlying system is commonly assumed to be mono- or multi-stable, and the system's response to 19–23.7 and 41 <inline-formula><mml:math id="M70" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> forcings is nonlinearly amplified at <inline-formula><mml:math id="M71" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M72" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> timescale <xref ref-type="bibr" rid="bib1.bibx72" id="paren.32"/>, for example, at the combination tone <inline-formula><mml:math id="M73" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">95</mml:mn><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">19</mml:mn><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">23.7</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M74" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">kyr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx42" id="paren.33"/>. Many studies, however, use the terms <italic>nonlinear response</italic> <xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx2" id="paren.34"/> or <italic>nonlinear amplification</italic> <xref ref-type="bibr" rid="bib1.bibx78" id="paren.35"/> to refer to cases compatible with the generalized notion of resonance. Other types of resonance have been discussed in relation to the <inline-formula><mml:math id="M75" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M76" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles, including linear resonance <xref ref-type="bibr" rid="bib1.bibx28 bib1.bibx23" id="paren.36"/>, stochastic resonance <xref ref-type="bibr" rid="bib1.bibx6 bib1.bibx55" id="paren.37"/>, coherence resonance <xref ref-type="bibr" rid="bib1.bibx61 bib1.bibx12" id="paren.38"/>, and vibrational resonance <xref ref-type="bibr" rid="bib1.bibx72" id="paren.39"/>.</p>
      <p id="d2e1152">Despite such differences in dynamical mechanisms and underlying system types, several ice age models with distinct approaches successfully simulate proxy records with similar accuracy, reproducing the <inline-formula><mml:math id="M77" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M78" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles. This raises an important question: if models with different mechanisms can reproduce the glacial cycles, what is the key factor that enables the <inline-formula><mml:math id="M79" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M80" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles, regardless of the specific mechanism? To address this question, we examine three previously proposed ice age models, each representing a different mechanism and underlying system type: one based on synchronization, one on resonance in a mono-stable system, and one on resonance in a multi-stable system with thresholds. Through sensitivity experiments changing the model's internal timescale and the amplitude of the forcing, we reveal that the key to enabling the 100 <inline-formula><mml:math id="M81" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles is the proximity of the intrinsic timescale of the underlying climate system to the <inline-formula><mml:math id="M82" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M83" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity of eccentricity cycles. Our results suggest that periodicity around <inline-formula><mml:math id="M84" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M85" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> occurs because of the match between the Earth system's intrinsic timescale and one astronomic timescale.</p>
      <p id="d2e1237">The remainder of this article is organized as follows. In Sect. 2 we present the three simple models of ice age cycles with different mechanisms for generating <inline-formula><mml:math id="M86" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M87" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles. In Sect. 3, we conduct sensitivity experiments by artificially varying the system's intrinsic timescale, demonstrating that it plays a crucial role in producing <inline-formula><mml:math id="M88" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M89" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles, regardless of the underlying mechanism. Section 4 is devoted to the discussion. In Sect. 5, we conclude the article.</p>
</sec>
<sec id="Ch1.S2">
  <label>2</label><title>Models for glacial cycles</title>
<sec id="Ch1.S2.SS1">
  <label>2.1</label><title>Self-sustained oscillator (SO) model representing the synchronization mechanism</title>
      <p id="d2e1291">A paradigmatic dynamical system featuring self-sustained oscillations is the oscillator of <xref ref-type="bibr" rid="bib1.bibx77" id="text.40"/>. Crucifix and colleagues have used the forced van der Pol oscillator as a mathematical model to investigate ice age dynamics <xref ref-type="bibr" rid="bib1.bibx17 bib1.bibx18 bib1.bibx21" id="paren.41"/>. We consider a generalized version of the model:

                <disp-formula specific-use="align" content-type="numbered"><mml:math id="M90" display="block"><mml:mtable displaystyle="true"><mml:mlabeledtr id="Ch1.E1"><mml:mtd><mml:mtext>1</mml:mtext></mml:mtd><mml:mtd><mml:mstyle class="stylechange" displaystyle="true"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mi>y</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">κ</mml:mi><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mi mathvariant="italic">μ</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:mfrac></mml:mstyle><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="Ch1.E2"><mml:mtd><mml:mtext>2</mml:mtext></mml:mtd><mml:mtd><mml:mstyle class="stylechange" displaystyle="true"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mover accent="true"><mml:mi>y</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">α</mml:mi><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:mi mathvariant="italic">β</mml:mi><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mo>-</mml:mo><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>-</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">ν</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">η</mml:mi><mml:msup><mml:mi>I</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mlabeledtr></mml:mtable></mml:math></disp-formula>

          with <inline-formula><mml:math id="M91" display="inline"><mml:mrow><mml:mi>y</mml:mi><mml:mo>=</mml:mo><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi><mml:mo>-</mml:mo><mml:mi mathvariant="italic">δ</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:math></inline-formula>, linking the variable <inline-formula><mml:math id="M92" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> with the ice volume proxy <inline-formula><mml:math id="M93" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> with an offset <inline-formula><mml:math id="M94" display="inline"><mml:mrow><mml:mi mathvariant="italic">δ</mml:mi><mml:mo>+</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:math></inline-formula>. Variable <inline-formula><mml:math id="M95" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> abstractly represents the “climate” state that determines whether the system is in the glaciation or the deglaciation phase, in combination with the insolation. It could represent the oceanic state <xref ref-type="bibr" rid="bib1.bibx21" id="paren.42"/>, the carbon cycle, dust concentrations, or their mixed effect. Variable <inline-formula><mml:math id="M96" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is the standardized summer solstice insolation anomaly at 65° N, and the model's parameters are denoted with Greek labels. The nonlinear term <inline-formula><mml:math id="M97" display="inline"><mml:mrow><mml:mi mathvariant="italic">η</mml:mi><mml:msup><mml:mi>I</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is included to take into account the lower sensitivity of the ice volume in the cold period <xref ref-type="bibr" rid="bib1.bibx59" id="paren.43"/>. Equations (<xref ref-type="disp-formula" rid="Ch1.E1"/>) and (<xref ref-type="disp-formula" rid="Ch1.E2"/>) contain not only the van der Pol equation but also the equation of <xref ref-type="bibr" rid="bib1.bibx24" id="text.44"/> due to the cubic term <inline-formula><mml:math id="M98" display="inline"><mml:mrow><mml:mi mathvariant="italic">β</mml:mi><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>, and the Hill equation <xref ref-type="bibr" rid="bib1.bibx46" id="paren.45"/> due to the multiplicative force <inline-formula><mml:math id="M99" display="inline"><mml:mrow><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>x</mml:mi><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> (see Appendix B for details). Thus, it is expected to have greater flexibility to accommodate complex nonlinear oscillations than the original forced van der Pol equation.</p>
      <p id="d2e1566">The parameters in Eqs. (<xref ref-type="disp-formula" rid="Ch1.E1"/>) and (<xref ref-type="disp-formula" rid="Ch1.E2"/>) and <inline-formula><mml:math id="M100" display="inline"><mml:mi mathvariant="italic">δ</mml:mi></mml:math></inline-formula> are tuned to minimize the mean squared errors between the simulated and observed <inline-formula><mml:math id="M101" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> records over the last 1 <inline-formula><mml:math id="M102" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula> (see Appendix B). The model reproduces the record of glacial cycles quite well (Fig. <xref ref-type="fig" rid="F2"/>b, pink; <inline-formula><mml:math id="M103" display="inline"><mml:mrow><mml:mi>R</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.88</mml:mn></mml:mrow></mml:math></inline-formula>) including the 95 <inline-formula><mml:math id="M104" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity (Fig. S2c in the Supplement).  For zero insolation anomaly <inline-formula><mml:math id="M105" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, the underlying system possesses self-sustained oscillations with a period of 91.7 <inline-formula><mml:math id="M106" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> (Fig. <xref ref-type="fig" rid="F2"/>b, sky blue). Such self-sustained oscillations occur over a range of insolation anomaly <inline-formula><mml:math id="M107" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.66</mml:mn><mml:mo>&lt;</mml:mo><mml:mi>I</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.075</mml:mn></mml:mrow></mml:math></inline-formula>. The oscillation period varies moderately over the range <inline-formula><mml:math id="M108" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.66</mml:mn><mml:mo>&lt;</mml:mo><mml:mi>I</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.075</mml:mn></mml:mrow></mml:math></inline-formula> with a mean of about 90 <inline-formula><mml:math id="M109" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> (Fig. S1 in the Supplement). The internal oscillations capture the slow buildup and the rapid disintegration of ice sheets. Under the astronomical forcing, the frequency entrainment occurs mainly at <inline-formula><mml:math id="M110" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">95</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M111" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">kyr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> near that of self-sustained oscillations, <inline-formula><mml:math id="M112" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">91.7</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M113" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">kyr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. Equations (<xref ref-type="disp-formula" rid="Ch1.E1"/>) and (<xref ref-type="disp-formula" rid="Ch1.E2"/>) are hereafter called the Self-sustained Oscillator (SO) model.</p>

      <fig id="F2" specific-use="star"><label>Figure 2</label><caption><p id="d2e1756">Forced and unforced simulations of glacial cycles over the last 1 <inline-formula><mml:math id="M114" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula>: <bold>(a)</bold> standardized summer solstice insolation at 65° N. <bold>(b)</bold> SO model with forcing (pink) and without forcing (light blue).  <bold>(c)</bold> VCV18 model with forcing (violet) and without forcing (light blue). <bold>(d)</bold> G24-3 model  with forcing (green) and without forcing (light blue). For all three models, the corresponding scaled versions of the paleoclimatic record are shown by the black dashed line.</p></caption>
          <graphic xlink:href="https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025-f02.png"/>

        </fig>

</sec>
<sec id="Ch1.S2.SS2">
  <label>2.2</label><title>Verbitsky–Crucifix–Volobuev model representing the resonance mechanism in monostable system</title>
      <p id="d2e1793"><xref ref-type="bibr" rid="bib1.bibx78" id="text.46"/> introduced a simple model of ice age cycles deduced from a scaling analysis of the governing physical laws (hereafter VCV18 model). The equations for the glaciation area (<inline-formula><mml:math id="M115" display="inline"><mml:mi>S</mml:mi></mml:math></inline-formula>), the basal temperature (<inline-formula><mml:math id="M116" display="inline"><mml:mi mathvariant="italic">θ</mml:mi></mml:math></inline-formula>), and the ocean temperature (<inline-formula><mml:math id="M117" display="inline"><mml:mi mathvariant="italic">ω</mml:mi></mml:math></inline-formula>) are given by

                <disp-formula specific-use="align"><mml:math id="M118" display="block"><mml:mtable displaystyle="true"><mml:mtr><mml:mtd><mml:mstyle class="stylechange" displaystyle="true"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mover accent="true"><mml:mi>S</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mn mathvariant="normal">4</mml:mn><mml:mn mathvariant="normal">5</mml:mn></mml:mfrac></mml:mstyle><mml:msup><mml:mi mathvariant="italic">ζ</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:msup><mml:mi>S</mml:mi><mml:mrow><mml:mn mathvariant="normal">3</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:msup><mml:mo>(</mml:mo><mml:mi>a</mml:mi><mml:mo>-</mml:mo><mml:mi mathvariant="italic">ε</mml:mi><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">κ</mml:mi><mml:mi mathvariant="italic">ω</mml:mi><mml:mo>-</mml:mo><mml:mi>c</mml:mi><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mstyle displaystyle="true" class="stylechange"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:mover accent="true"><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:msup><mml:mi mathvariant="italic">ζ</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:msup><mml:mi>S</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:msup><mml:mo>(</mml:mo><mml:mi>a</mml:mi><mml:mo>-</mml:mo><mml:mi mathvariant="italic">ε</mml:mi><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">κ</mml:mi><mml:mi mathvariant="italic">ω</mml:mi><mml:mo>)</mml:mo><mml:mo mathvariant="italic">{</mml:mo><mml:mi mathvariant="italic">α</mml:mi><mml:mi mathvariant="italic">ω</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">β</mml:mi><mml:mo>[</mml:mo><mml:mi>S</mml:mi><mml:mo>-</mml:mo><mml:msub><mml:mi>S</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>]</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">θ</mml:mi><mml:mo mathvariant="italic">}</mml:mo></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mstyle displaystyle="true" class="stylechange"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mover accent="true"><mml:mi mathvariant="italic">ω</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:msub><mml:mi mathvariant="italic">γ</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">γ</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>[</mml:mo><mml:mi>S</mml:mi><mml:mo>-</mml:mo><mml:msub><mml:mi>S</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>]</mml:mo><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">γ</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msub><mml:mi mathvariant="italic">ω</mml:mi><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:math></disp-formula>

          where <inline-formula><mml:math id="M119" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is the standardized summer solstice insolation at 65° N. The ice volume is given as <inline-formula><mml:math id="M120" display="inline"><mml:mrow><mml:mi>V</mml:mi><mml:mo>=</mml:mo><mml:mi mathvariant="italic">ζ</mml:mi><mml:msup><mml:mi>S</mml:mi><mml:mrow><mml:mn mathvariant="normal">5</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. See Table 1 in <xref ref-type="bibr" rid="bib1.bibx78" id="text.47"/> for the parameter values. Since the system becomes numerically unstable near <inline-formula><mml:math id="M121" display="inline"><mml:mrow><mml:mi>S</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, we reset the <inline-formula><mml:math id="M122" display="inline"><mml:mi>S</mml:mi></mml:math></inline-formula> value to <inline-formula><mml:math id="M123" display="inline"><mml:mrow><mml:msup><mml:mn mathvariant="normal">10</mml:mn><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> if it falls below <inline-formula><mml:math id="M124" display="inline"><mml:mrow><mml:msup><mml:mn mathvariant="normal">10</mml:mn><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. The VCV18 model can roughly simulate changes in sea level as shown in Fig. <xref ref-type="fig" rid="F2"/>c (violet). Although the simulated sea level does not  capture the amplitude and timing of all deglaciations (specifically, the last one), the model exhibits prominent <inline-formula><mml:math id="M125" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M126" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> power consistently with the record (Fig. S3c in the Supplement). In the absence of forcing (<inline-formula><mml:math id="M127" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>≡</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>), the system has a single stable equilibrium whose Jacobian matrix has one real negative eigenvalue and a pair of complex conjugate eigenvalues with negative real parts. The imaginary part of the latter pair defines the <italic>natural frequency</italic> of the system. In this article, we refer to the inverse of the natural frequency as the <italic>natural period</italic>. Under the standard parameter set, this natural period is 95 <inline-formula><mml:math id="M128" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>.</p>
      <p id="d2e2172">Although the astronomical forcing has most of its power at <inline-formula><mml:math id="M129" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">20</mml:mn></mml:mrow></mml:math></inline-formula> and 41 <inline-formula><mml:math id="M130" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> bands, the dominant power in the climate response is concentrated near <inline-formula><mml:math id="M131" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M132" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. Since the system does not exhibit self-sustained oscillations, the appearance of <inline-formula><mml:math id="M133" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M134" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles in the VCV18 model cannot be qualified as a phenomenon of synchronization. Instead, it must be related to a nonlinear amplification of the response <xref ref-type="bibr" rid="bib1.bibx78" id="paren.48"/>, i.e., nonlinear resonance, as shown in Sect. 3.</p>
</sec>
<sec id="Ch1.S2.SS3">
  <label>2.3</label><title>Ganopolski model representing the resonance mechanism in multistable systems</title>
      <p id="d2e2241"><xref ref-type="bibr" rid="bib1.bibx25" id="text.49"/> discusses three simple models of ice age cycles in his Generalized Milankovitch Theory (hereafter G24-1,2,3). The G24-3 model is a model derived from ice age simulations using the Earth system model of intermediate complexity CLIMBER-2 <xref ref-type="bibr" rid="bib1.bibx13 bib1.bibx26 bib1.bibx80" id="paren.50"/>. The change in ice volume <inline-formula><mml:math id="M135" display="inline"><mml:mi>v</mml:mi></mml:math></inline-formula> is defined in the glaciation and deglaciation regimes, respectively, as

                <disp-formula id="Ch1.Ex4"><mml:math id="M136" display="block"><mml:mstyle class="stylechange" displaystyle="true"/><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mover accent="true"><mml:mi>v</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mfenced open="{" close=""><mml:mtable columnspacing="1em" class="cases" rowspacing="0.2ex" columnalign="left left" framespacing="0em"><mml:mtr><mml:mtd><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">e</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:mi>v</mml:mi></mml:mrow><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if </mml:mtext><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mtext> (glaciation regime)</mml:mtext></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:mo>-</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>v</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if </mml:mtext><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">2</mml:mn><mml:mtext> (deglaciation regime)</mml:mtext><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:mfenced></mml:mrow></mml:math></disp-formula>

          where <inline-formula><mml:math id="M137" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">30</mml:mn></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M138" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">10</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M139" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> are relaxation timescales in each regime estimated from CLIMBER-2 experiments <xref ref-type="bibr" rid="bib1.bibx13" id="paren.51"/>. The term <inline-formula><mml:math id="M140" display="inline"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">e</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> represents either of two stable equilibria depending on the state <inline-formula><mml:math id="M141" display="inline"><mml:mi>v</mml:mi></mml:math></inline-formula> and the 65° N summer solstice insolation anomaly <inline-formula><mml:math id="M142" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> relative its mean over the last 1 <inline-formula><mml:math id="M143" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula>:

                <disp-formula id="Ch1.Ex5"><mml:math id="M144" display="block"><mml:mstyle class="stylechange" displaystyle="true"/><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">e</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mfenced open="{" close=""><mml:mtable class="cases" columnspacing="1em" rowspacing="0.2ex" columnalign="left left" framespacing="0em"><mml:mtr><mml:mtd><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if </mml:mtext><mml:mi>f</mml:mi><mml:mo>&lt;</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mtext>, or </mml:mtext><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>&lt;</mml:mo><mml:mi>f</mml:mi><mml:mo>&lt;</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mtext> and </mml:mtext><mml:mi>v</mml:mi><mml:mo>&gt;</mml:mo><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">u</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">i</mml:mi></mml:msub></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if </mml:mtext><mml:mi>f</mml:mi><mml:mo>&gt;</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mtext>, or </mml:mtext><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>&lt;</mml:mo><mml:mi>f</mml:mi><mml:mo>&lt;</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mtext> and </mml:mtext><mml:mi>v</mml:mi><mml:mo>&lt;</mml:mo><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">u</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:mfenced></mml:mrow></mml:math></disp-formula>

          where <inline-formula><mml:math id="M145" display="inline"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>≤</mml:mo><mml:mi>f</mml:mi><mml:mo>≤</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> is the range of multiple equilibria, with <inline-formula><mml:math id="M146" display="inline"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">16</mml:mn></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M147" display="inline"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">16</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M148" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">Wm</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. The function <inline-formula><mml:math id="M149" display="inline"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>+</mml:mo><mml:msqrt><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>-</mml:mo><mml:mi>f</mml:mi></mml:mrow><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:msqrt></mml:mrow></mml:math></inline-formula> represents the glacial equilibrium, and <inline-formula><mml:math id="M150" display="inline"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">i</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula> is the interglacial equilibrium. The unstable equilibrium separating the glacial and interglacial basins is given by <inline-formula><mml:math id="M151" display="inline"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">u</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msqrt><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>-</mml:mo><mml:mi>f</mml:mi></mml:mrow><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:msqrt></mml:mrow></mml:math></inline-formula> (see Fig. 4 in <xref ref-type="bibr" rid="bib1.bibx25" id="altparen.52"/>). Note that <inline-formula><mml:math id="M152" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is an anomaly and not scaled by its standard deviation, different to <inline-formula><mml:math id="M153" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> in the previous two models. The transition from the glaciation regime (<inline-formula><mml:math id="M154" display="inline"><mml:mrow><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>) to the deglaciation regime (<inline-formula><mml:math id="M155" display="inline"><mml:mrow><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:math></inline-formula>) occurs if three conditions are met: <inline-formula><mml:math id="M156" display="inline"><mml:mrow><mml:mi>v</mml:mi><mml:mo>&gt;</mml:mo><mml:msub><mml:mi>v</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M157" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M158" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>f</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M159" display="inline"><mml:mrow><mml:msub><mml:mi>v</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1.4</mml:mn><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is the critical ice volume, above which the ice sheets are likely to collapse. The transition from the deglaciation regime (<inline-formula><mml:math id="M160" display="inline"><mml:mrow><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:math></inline-formula>) to the glaciation regime (<inline-formula><mml:math id="M161" display="inline"><mml:mrow><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>) occurs if <inline-formula><mml:math id="M162" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> drops below <inline-formula><mml:math id="M163" display="inline"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>. Since <inline-formula><mml:math id="M164" display="inline"><mml:mi>v</mml:mi></mml:math></inline-formula> should remain non-negative, we reset it to 0 at each integration time step if it becomes negative.</p>
      <p id="d2e2915">The G24-3 model simulates the glacial cycles well (<inline-formula><mml:math id="M165" display="inline"><mml:mrow><mml:mi>R</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.82</mml:mn></mml:mrow></mml:math></inline-formula> over 1 <inline-formula><mml:math id="M166" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula>) and has two stable equilibria for <inline-formula><mml:math id="M167" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula> (Fig. <xref ref-type="fig" rid="F2"/>d). By construction, the G24-3 model  does not produce self-sustained oscillations for constant insolation because its regime transitions require threshold crossings in insolation. <xref ref-type="bibr" rid="bib1.bibx25" id="text.53"/> mentions that the characteristic timescales of the model are <inline-formula><mml:math id="M168" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">30</mml:mn></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M169" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">10</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M170" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, and the model has no intrinsic timescale close to 100 <inline-formula><mml:math id="M171" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. However, the intrinsic timescale of the G24-3 model may be considered much longer than the relaxation times <inline-formula><mml:math id="M172" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">30</mml:mn></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M173" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">10</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M174" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. First, assuming the average insolation <inline-formula><mml:math id="M175" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, the time in which the ice volume increases from <inline-formula><mml:math id="M176" display="inline"><mml:mrow><mml:mi>v</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula> to the critical ice volume <inline-formula><mml:math id="M177" display="inline"><mml:mrow><mml:msub><mml:mi>v</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is <inline-formula><mml:math id="M178" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mi>ln⁡</mml:mi><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:msub><mml:mi>v</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>≈</mml:mo><mml:mn mathvariant="normal">51.5</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M179" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. Even after the ice volume exceeds <inline-formula><mml:math id="M180" display="inline"><mml:mrow><mml:msub><mml:mi>v</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, the ice sheets continue to grow until the insolation anomaly <inline-formula><mml:math id="M181" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> changes from negative to positive. While this time lag varies depending on the phase of the precession cycles, half of the precession period, approximately 10 <inline-formula><mml:math id="M182" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, is a reasonable expected value. Adding this lag on top of 51.5 <inline-formula><mml:math id="M183" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, the total period from glacial inception to the onset of deglaciation is estimated as 61.5 <inline-formula><mml:math id="M184" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. Second, the time it takes for the ice to melt is about <inline-formula><mml:math id="M185" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">10</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M186" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. After this period of deglaciation, which usually continues during <inline-formula><mml:math id="M187" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, the system waits for glacial inception triggered by the drop in <inline-formula><mml:math id="M188" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> below <inline-formula><mml:math id="M189" display="inline"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>. This waiting time is roughly <inline-formula><mml:math id="M190" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:math></inline-formula> precession cycle, i.e., <inline-formula><mml:math id="M191" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">5</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M192" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. The sum of the glaciation timescale and the deglaciation timescale for G24-3 model is <inline-formula><mml:math id="M193" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mi>ln⁡</mml:mi><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1.5</mml:mn></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>+</mml:mo><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">61.5</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M194" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, while the timescale to complete a cycle including the time lags is <inline-formula><mml:math id="M195" display="inline"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mi>ln⁡</mml:mi><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1.5</mml:mn></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>+</mml:mo><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mo>+</mml:mo><mml:mn mathvariant="normal">15</mml:mn><mml:mo>=</mml:mo><mml:mn mathvariant="normal">76.5</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M196" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. This timescale is closer to the 95 <inline-formula><mml:math id="M197" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity period than other fundamental astronomical periods.</p>
</sec>
</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Sensitivity experiments</title>
      <p id="d2e3407">In the previous section, we showed that the three models of ice age cycles exhibit distinct types of underlying dynamics, each having a characteristic timescale: the period of a self-sustained oscillation, the period corresponding to the natural frequency (i.e., natural period), and the timescale for forming a cycle. These system's intrinsic timescales are schematically illustrated in Fig. <xref ref-type="fig" rid="F3"/>. Here, we conduct sensitivity experiments on the models described in Sect. 2 to demonstrate that an intrinsic timescale close to <inline-formula><mml:math id="M198" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M199" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> is the key to enabling a periodicity around 100 <inline-formula><mml:math id="M200" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> in all three types of models.</p>

      <fig id="F3"><label>Figure 3</label><caption><p id="d2e3440">Three types of proposed underlying dynamics of ice age cycles and their characteristic timescales. <bold>(a)</bold> Self-sustained oscillation with a period. <bold>(b)</bold> Damped oscillation characterized by a period corresponding to the natural frequency. <bold>(c)</bold> A bistable system characterized by a timescale for forming a cycle, which includes the timescale of glaciation, the timescale of deglaciation, the time lag before glaciation is triggered by insolation, and the time lag before deglaciation is triggered.</p></caption>
        <graphic xlink:href="https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025-f03.png"/>

      </fig>

<sec id="Ch1.S3.SS1">
  <label>3.1</label><title>Responses to the astronomical forcing</title>
      <p id="d2e3465">First, we show that the three models exhibit different responses to astronomical forcing. The models are run with a scaled insolation forcing: <inline-formula><mml:math id="M201" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>I</mml:mi><mml:mo stretchy="false" mathvariant="normal">̃</mml:mo></mml:mover><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mi>A</mml:mi><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> in the SO model and the VCV18 model and <inline-formula><mml:math id="M202" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>f</mml:mi><mml:mo mathvariant="normal" stretchy="false">̃</mml:mo></mml:mover><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mi>A</mml:mi><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> in the G24-3 model. The original simulations correspond to <inline-formula><mml:math id="M203" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>. The changes in the PSD for varying <inline-formula><mml:math id="M204" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> in steps of 0.02 are shown in Fig. <xref ref-type="fig" rid="F4"/>. Specific time series and PSDs for <inline-formula><mml:math id="M205" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, 0.5, 1, 1.5, and 2 are shown in Figs. S2–S4 in the Supplement.</p>

      <fig id="F4" specific-use="star"><label>Figure 4</label><caption><p id="d2e3562">Power spectral  density (PSD) for different amplitudes <inline-formula><mml:math id="M206" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> of the astronomical forcing: <bold>(a)</bold> SO model. <bold>(b)</bold> VCV18 model. <bold>(c)</bold> G24-3 model. The PSDs are obtained from simulations over the last 1 <inline-formula><mml:math id="M207" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula>. The magenta dashed lines indicate the major astronomical frequencies (the numbers show the corresponding periods). The precession band, 19–23 <inline-formula><mml:math id="M208" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, is not shown since its power is comparatively minor.</p></caption>
          <graphic xlink:href="https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025-f04.png"/>

        </fig>

      <p id="d2e3604">In the SO model, as shown in Fig. <xref ref-type="fig" rid="F4"/>a, the PSD has its maximum at 91.7 <inline-formula><mml:math id="M209" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> for zero forcing amplitude, <inline-formula><mml:math id="M210" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, corresponding to self-sustained oscillations. For small <inline-formula><mml:math id="M211" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">0.84</mml:mn></mml:mrow></mml:math></inline-formula>, frequency locking to a major astronomical period is not achieved (Fig. S2a and b in the Supplement). The frequency locking to a 82 <inline-formula><mml:math id="M212" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> double-obliquity period is realized for a very narrow range <inline-formula><mml:math id="M213" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.86</mml:mn><mml:mo>≤</mml:mo><mml:mi>A</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">0.88</mml:mn></mml:mrow></mml:math></inline-formula>. The frequency locking at 95 <inline-formula><mml:math id="M214" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> occurs for a wide range of <inline-formula><mml:math id="M215" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.90</mml:mn><mml:mo>≤</mml:mo><mml:mi>A</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">1.52</mml:mn></mml:mrow></mml:math></inline-formula>. For larger <inline-formula><mml:math id="M216" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula>, the principal period shifts toward the larger side, exhibiting frequency lockings to 124 or 405 <inline-formula><mml:math id="M217" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>.</p>
      <p id="d2e3706">In the VCV18 model shown in Fig. <xref ref-type="fig" rid="F4"/>b, the total power is quite small for low <inline-formula><mml:math id="M218" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> since the underlying dynamics is a damped oscillation. For <inline-formula><mml:math id="M219" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> less than half of the original value, the PSD has a maximum at 41 <inline-formula><mml:math id="M220" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, although it is too small to be clearly seen in Fig. <xref ref-type="fig" rid="F4"/>b. This is simply the linear response to the 41 <inline-formula><mml:math id="M221" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> obliquity cycles. A large power appears at the 95 <inline-formula><mml:math id="M222" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity period as <inline-formula><mml:math id="M223" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> increases to more than 0.5. This resonance with 95 <inline-formula><mml:math id="M224" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity cycles is actually a nonlinear resonance to the combination tone between 19–23.7 <inline-formula><mml:math id="M225" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> precession cycles. This nonlinear resonance is found near the system's natural period of 95 <inline-formula><mml:math id="M226" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. This is in line with the classical notion that the resonance typically occurs if the frequency of external forcing matches the natural frequency of the system.</p>
      <p id="d2e3783">In the G24-3 model shown in Fig. <xref ref-type="fig" rid="F4"/>c, the power is zero for low forcing amplitude <inline-formula><mml:math id="M227" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">0.36</mml:mn></mml:mrow></mml:math></inline-formula> because glacial inception cannot be triggered. Glacial inception is possible for <inline-formula><mml:math id="M228" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>≥</mml:mo><mml:mn mathvariant="normal">0.38</mml:mn></mml:mrow></mml:math></inline-formula>. The main peak is located at 405 <inline-formula><mml:math id="M229" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> for <inline-formula><mml:math id="M230" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.38</mml:mn><mml:mo>≤</mml:mo><mml:mi>A</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">0.42</mml:mn></mml:mrow></mml:math></inline-formula>, at 124 <inline-formula><mml:math id="M231" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> for <inline-formula><mml:math id="M232" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.44</mml:mn><mml:mo>≤</mml:mo><mml:mi>A</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">0.54</mml:mn></mml:mrow></mml:math></inline-formula>, and at 95 <inline-formula><mml:math id="M233" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> for the wide range of <inline-formula><mml:math id="M234" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.56</mml:mn><mml:mo>≤</mml:mo><mml:mi>A</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">1.42</mml:mn></mml:mrow></mml:math></inline-formula>. This occurs because the frequency of threshold crossing increases as <inline-formula><mml:math id="M235" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> increases. The principal period remains close to 100 <inline-formula><mml:math id="M236" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> for larger <inline-formula><mml:math id="M237" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>(</mml:mo><mml:mo>≥</mml:mo><mml:mn mathvariant="normal">1.44</mml:mn><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, but it is different from any major astronomical period.</p>
</sec>
<sec id="Ch1.S3.SS2">
  <label>3.2</label><title>Intrinsic timescales and responses</title>
      <p id="d2e3928">Next, we investigate the relationship between the principal period of the output and the intrinsic timescale of the model. For this purpose, we introduce a parameter <inline-formula><mml:math id="M238" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula> that modulates the timescale of the model, following previous studies <xref ref-type="bibr" rid="bib1.bibx21 bib1.bibx18" id="paren.54"/>. Each dynamical equation is scaled as <inline-formula><mml:math id="M239" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:mi>X</mml:mi></mml:mrow><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:mi>t</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:mtext>r.h.s</mml:mtext></mml:mrow></mml:math></inline-formula>. The larger <inline-formula><mml:math id="M240" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula>, the slower the temporal variation of the model variables. In the SO model, the period of self-sustained oscillations (originally <inline-formula><mml:math id="M241" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">91.7</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M242" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>) is scaled as <inline-formula><mml:math id="M243" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>. Similarly, in the VCV18 model, the natural period of the damped oscillations (originally <inline-formula><mml:math id="M244" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">95</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M245" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>) becomes <inline-formula><mml:math id="M246" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>. In the G24-3 model, the intrinsic timescale for the glaciation and deglaciation is scaled as <inline-formula><mml:math id="M247" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:mfenced open="(" close=")"><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mi>ln⁡</mml:mi><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:msub><mml:mi>v</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>+</mml:mo><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:mfenced><mml:mo>=</mml:mo><mml:mn mathvariant="normal">61.5</mml:mn><mml:mi>r</mml:mi></mml:mrow></mml:math></inline-formula> kyr. Adding the time lags required for the astronomical conditions to be met, the timescale for forming a cycle is <inline-formula><mml:math id="M248" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mi>r</mml:mi><mml:mfenced close=")" open="("><mml:mrow><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mi>ln⁡</mml:mi><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:msub><mml:mi>V</mml:mi><mml:mi mathvariant="normal">g</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:msub><mml:mi>v</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>+</mml:mo><mml:msub><mml:mi>t</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:mfenced><mml:mo>+</mml:mo><mml:mn mathvariant="normal">15</mml:mn><mml:mo>=</mml:mo><mml:mn mathvariant="normal">61.5</mml:mn><mml:mi>r</mml:mi><mml:mo>+</mml:mo><mml:mn mathvariant="normal">15</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M249" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. The tempo of orbital forcing remains unchanged.</p>
      <p id="d2e4202">We run each model by varying <inline-formula><mml:math id="M250" display="inline"><mml:mrow><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>,</mml:mo><mml:mi>A</mml:mi><mml:mo>)</mml:mo><mml:mo>∈</mml:mo><mml:mo>[</mml:mo><mml:mn mathvariant="normal">0.5</mml:mn><mml:mo>,</mml:mo><mml:mn mathvariant="normal">1.5</mml:mn><mml:mo>]</mml:mo><mml:mo>×</mml:mo><mml:mo>[</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>,</mml:mo><mml:mn mathvariant="normal">2</mml:mn><mml:mo>]</mml:mo></mml:mrow></mml:math></inline-formula> and measure the principal period of the simulated ice age cycles from the PSD <inline-formula><mml:math id="M251" display="inline"><mml:mrow><mml:mi>S</mml:mi><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> as <inline-formula><mml:math id="M252" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">P</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:msub><mml:mtext>argmax</mml:mtext><mml:mi>f</mml:mi></mml:msub><mml:mi>S</mml:mi><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>. We judge that the measured principal period <inline-formula><mml:math id="M253" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">P</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is virtually identical to one of the major astronomical periods, <inline-formula><mml:math id="M254" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, if <inline-formula><mml:math id="M255" display="inline"><mml:mrow><mml:mo>|</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">P</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>|</mml:mo><mml:mo>&lt;</mml:mo><mml:mi mathvariant="italic">ϵ</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M256" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">19</mml:mn></mml:mrow></mml:math></inline-formula>, 22.4, 23.7, 41, 82, 95, 124, or 405 <inline-formula><mml:math id="M257" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> (note that 82 <inline-formula><mml:math id="M258" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> corresponds to twice the obliquity cycle). The parameter <inline-formula><mml:math id="M259" display="inline"><mml:mi mathvariant="italic">ϵ</mml:mi></mml:math></inline-formula> is set to be small, specifically <inline-formula><mml:math id="M260" display="inline"><mml:mrow><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.028</mml:mn></mml:mrow></mml:math></inline-formula> for the SO model and <inline-formula><mml:math id="M261" display="inline"><mml:mrow><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.04</mml:mn></mml:mrow></mml:math></inline-formula> for the VCV18 and G24-3 models. Only for the case <inline-formula><mml:math id="M262" display="inline"><mml:mrow><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.04</mml:mn></mml:mrow></mml:math></inline-formula>, some <inline-formula><mml:math id="M263" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">P</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> can satisfy the condition <inline-formula><mml:math id="M264" display="inline"><mml:mrow><mml:mo>|</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">P</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>|</mml:mo><mml:mo>&lt;</mml:mo><mml:mi mathvariant="italic">ϵ</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> for both <inline-formula><mml:math id="M265" display="inline"><mml:mrow><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">22.5</mml:mn></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M266" display="inline"><mml:mrow><mml:msub><mml:mi>P</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">23.7</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M267" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> simultaneously; in such a case, we choose the closer one to be the simulated principal period. The results are shown in Fig. <xref ref-type="fig" rid="F5"/>, as will be explained later.</p>
      <p id="d2e4502">We also calculate a measure of resonance, specifically the response amplitude of signal <inline-formula><mml:math id="M268" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> at a given frequency <inline-formula><mml:math id="M269" display="inline"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (i.e., period <inline-formula><mml:math id="M270" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:msub><mml:mi>f</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>): <inline-formula><mml:math id="M271" display="inline"><mml:mrow><mml:mi>Q</mml:mi><mml:mo>=</mml:mo><mml:msqrt><mml:mrow><mml:msubsup><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">s</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msubsup><mml:mo>+</mml:mo><mml:msubsup><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">c</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msubsup></mml:mrow></mml:msqrt></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M272" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">2</mml:mn><mml:mrow><mml:mi>n</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:msubsup><mml:mo>∫</mml:mo><mml:mrow><mml:mo>-</mml:mo><mml:mi>n</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow><mml:mn mathvariant="normal">0</mml:mn></mml:msubsup><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mi>sin⁡</mml:mi><mml:mo>(</mml:mo><mml:mn mathvariant="normal">2</mml:mn><mml:mi mathvariant="italic">π</mml:mi><mml:mi>t</mml:mi><mml:mo>/</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mi mathvariant="normal">d</mml:mi><mml:mi>t</mml:mi></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M273" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">2</mml:mn><mml:mrow><mml:mi>n</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:msubsup><mml:mo>∫</mml:mo><mml:mrow><mml:mo>-</mml:mo><mml:mi>n</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow><mml:mn mathvariant="normal">0</mml:mn></mml:msubsup><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mi>cos⁡</mml:mi><mml:mo>(</mml:mo><mml:mn mathvariant="normal">2</mml:mn><mml:mi mathvariant="italic">π</mml:mi><mml:mi>t</mml:mi><mml:mo>/</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mi mathvariant="normal">d</mml:mi><mml:mi>t</mml:mi></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M274" display="inline"><mml:mi>n</mml:mi></mml:math></inline-formula> is chosen so that the integration interval spans at most the last 1000 <inline-formula><mml:math id="M275" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, that is <inline-formula><mml:math id="M276" display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo><mml:mfenced open="⌊" close="⌋"><mml:mrow><mml:mn mathvariant="normal">1000</mml:mn><mml:mo>/</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:mfenced></mml:mrow></mml:math></inline-formula>. Since the parameter <inline-formula><mml:math id="M277" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> is related to the PSD as <inline-formula><mml:math id="M278" display="inline"><mml:mrow><mml:mi>S</mml:mi><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo><mml:mo>∝</mml:mo><mml:msup><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, the resonance can also be quantified by the PSD <inline-formula><mml:math id="M279" display="inline"><mml:mrow><mml:mi>S</mml:mi><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>. However, here we employ <inline-formula><mml:math id="M280" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> as it is a widely accepted measure of resonance <xref ref-type="bibr" rid="bib1.bibx72 bib1.bibx65" id="paren.55"/>. The results are shown in Fig. <xref ref-type="fig" rid="F6"/>.</p>
      <p id="d2e4822">In the SO model shown in Fig. <xref ref-type="fig" rid="F5"/>a, we identify regions where the principal period aligns with one of the major astronomical cycles. Each region originates from a point along the horizontal axis where the scaled internal period, <inline-formula><mml:math id="M281" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>, matches a major astronomical period. These regions resemble the so-called <italic>Arnold tongues</italic> observed in periodically forced systems <xref ref-type="bibr" rid="bib1.bibx63" id="paren.56"/>. Within an Arnold tongue, the mean oscillation frequency – defined as the number of cycles over a long time interval – is locked to a forcing frequency or its simple rational multiple. However, the regions in Fig. <xref ref-type="fig" rid="F5"/>a do not strictly qualify as Arnold tongues because the principal frequency identified by the maximum peak of the PSD does not necessarily coincide with the mean oscillation frequency. We therefore refer to these as <italic>quasi-Arnold tongues</italic>: triangular regions where the principal frequency of a self-sustained oscillator under external forcing matches one of the forcing frequencies or a linear combination thereof (Fig. <xref ref-type="fig" rid="F5"/>a). Unlike the strict Arnold tongue, this concept only relies on the match between the principal and forcing frequencies, making it a more relaxed criterion. Notably, the quasi-Arnold tongue corresponding to the 95 <inline-formula><mml:math id="M282" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity is narrow and vertical, indicating that in the SO model, the 95 <inline-formula><mml:math id="M283" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycle reflects the system's intrinsic frequency.</p>

      <fig id="F5" specific-use="star"><label>Figure 5</label><caption><p id="d2e4873">Regime diagram in <inline-formula><mml:math id="M284" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula>–<inline-formula><mml:math id="M285" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula> space. <bold>(a)</bold> SO model (synchronization mechanism). <bold>(b)</bold> VCV18 model (nonlinear resonance mechanism in a damped oscillatory system). <bold>(c)</bold> G24-3 model (nonlinear resonance mechanism in a bistable system with thresholds). The principal period of the simulated dynamics is shown by the symbols in the legend. The most realistic simulations are obtained at <inline-formula><mml:math id="M286" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M287" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula> (black diamond). <inline-formula><mml:math id="M288" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> is the forcing amplitude. <inline-formula><mml:math id="M289" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula> is the parameter controlling the timescale of the underlying system. <inline-formula><mml:math id="M290" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> is the scaled intrinsic timescale in the SO model and the VCV18 model. In the G24-3 model, the scaled timescale for forming a cycle is <inline-formula><mml:math id="M291" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">61.5</mml:mn><mml:mi>r</mml:mi><mml:mo>+</mml:mo><mml:mn mathvariant="normal">15</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M292" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>.</p></caption>
          <graphic xlink:href="https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025-f05.png"/>

        </fig>

      <p id="d2e4993">The VCV18 model does not exhibit quasi-Arnold tongues that touch the horizontal axis at a single point (Fig. <xref ref-type="fig" rid="F5"/>b). For small but nonzero values of the forcing amplitude <inline-formula><mml:math id="M293" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula>, the principal period is 23.7 <inline-formula><mml:math id="M294" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> when the scaled natural period <inline-formula><mml:math id="M295" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> is less than roughly 41 and 41 <inline-formula><mml:math id="M296" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> when <inline-formula><mml:math id="M297" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>≳</mml:mo><mml:mn mathvariant="normal">41</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M298" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M299" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.5</mml:mn><mml:mo>≲</mml:mo><mml:mi>r</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:math></inline-formula>). These correspond to linear responses to the 23 <inline-formula><mml:math id="M300" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> precession and 41 <inline-formula><mml:math id="M301" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> obliquity components of the insolation forcing, respectively. For forcing amplitudes <inline-formula><mml:math id="M302" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> roughly between 0.5–1.5, three regions appear in succession as the natural period <inline-formula><mml:math id="M303" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> varies, each corresponding to a dominant period of 41, 95, and 124 <inline-formula><mml:math id="M304" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, respectively (Fig. <xref ref-type="fig" rid="F5"/>b). In each region, the corresponding <inline-formula><mml:math id="M305" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> parameter reaches its maximum, confirming that these are resonance phenomena (Fig. <xref ref-type="fig" rid="F6"/>b). Moreover, the 95 and 124 <inline-formula><mml:math id="M306" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> resonance regions incline toward larger values of <inline-formula><mml:math id="M307" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> as <inline-formula><mml:math id="M308" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> increases, respectively (Fig. <xref ref-type="fig" rid="F5"/>b). Such a shift in the natural period <inline-formula><mml:math id="M309" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> that yields the maximum amplitude is characteristic of nonlinear resonance <xref ref-type="bibr" rid="bib1.bibx65 bib1.bibx47" id="paren.57"/>. Near the realistic forcing amplitude of <inline-formula><mml:math id="M310" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>≃</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>, resonances at 41, 95, and 124 <inline-formula><mml:math id="M311" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> emerge when the scaled natural period <inline-formula><mml:math id="M312" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> approaches these respective timescales (Figs. <xref ref-type="fig" rid="F5"/>b and <xref ref-type="fig" rid="F6"/>b). This correspondence indicates that resonance is driven by a timescale match between the system's natural period and an astronomical period, in line with the classical concept of resonance. We therefore conclude that the proximity between the system's intrinsic timescale and the 95 <inline-formula><mml:math id="M313" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity period is crucial for producing the 95 <inline-formula><mml:math id="M314" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles in the VCV18 model as well. Note that the close numerical match between the natural period <inline-formula><mml:math id="M315" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">95</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M316" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> and the 95 <inline-formula><mml:math id="M317" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity period is purely coincidental, and the resonance at 95 <inline-formula><mml:math id="M318" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> can occur for a range of natural periods, <inline-formula><mml:math id="M319" display="inline"><mml:mrow><mml:mn mathvariant="normal">83</mml:mn><mml:mo>≤</mml:mo><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">118</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M320" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, for the realistic astronomical forcing <inline-formula><mml:math id="M321" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>.</p>

      <fig id="F6" specific-use="star"><label>Figure 6</label><caption><p id="d2e5317"><inline-formula><mml:math id="M322" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> spectrum as a function of the timescale control parameter <inline-formula><mml:math id="M323" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula>.  <bold>(a)</bold> SO model (synchronization mechanism). <bold>(b)</bold> VCV18 model (nonlinear resonance mechanism in a damped oscillatory system). <bold>(c)</bold> G24-3 model (nonlinear resonance mechanism in a bistable system with thresholds). The most realistic simulations are obtained at <inline-formula><mml:math id="M324" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M325" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">95</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> for the 95 <inline-formula><mml:math id="M326" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> period is maximal. <inline-formula><mml:math id="M327" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>T</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> is the scaled intrinsic timescale in the SO model and the VCV18 model. In the G24-3 model, the scaled timescale for forming a cycle is <inline-formula><mml:math id="M328" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">61.5</mml:mn><mml:mi>r</mml:mi><mml:mo>+</mml:mo><mml:mn mathvariant="normal">15</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M329" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>.</p></caption>
          <graphic xlink:href="https://esd.copernicus.org/articles/16/1569/2025/esd-16-1569-2025-f06.png"/>

        </fig>

      <p id="d2e5428">In the G24-3 model, as shown in Fig. <xref ref-type="fig" rid="F5"/>c, 124 <inline-formula><mml:math id="M330" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> glacial cycles as well as 405 <inline-formula><mml:math id="M331" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles occur for a wide range of <inline-formula><mml:math id="M332" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula>, i.e., virtually regardless of the scaled timescale for forming a cycle, <inline-formula><mml:math id="M333" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>. The 95 <inline-formula><mml:math id="M334" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles also occur for a wide range of scaled timescales <inline-formula><mml:math id="M335" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> if <inline-formula><mml:math id="M336" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> is around 0.6. However, the range of <inline-formula><mml:math id="M337" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> giving the 95 <inline-formula><mml:math id="M338" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles is limited to <inline-formula><mml:math id="M339" display="inline"><mml:mrow><mml:mn mathvariant="normal">66</mml:mn><mml:mo>≲</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>r</mml:mi><mml:mo>)</mml:mo><mml:mo>≲</mml:mo><mml:mn mathvariant="normal">90</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M340" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> at the realistic forcing amplitude <inline-formula><mml:math id="M341" display="inline"><mml:mrow><mml:mi>A</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>. Thus, also in the G24-3 model, the intrinsic timescale is key to having the 95 <inline-formula><mml:math id="M342" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles. We calculate the <inline-formula><mml:math id="M343" display="inline"><mml:mi>Q</mml:mi></mml:math></inline-formula> parameters also for this model (Fig. <xref ref-type="fig" rid="F6"/>c). Among others, <inline-formula><mml:math id="M344" display="inline"><mml:mrow><mml:msub><mml:mi>Q</mml:mi><mml:mn mathvariant="normal">95</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> corresponding to 95 <inline-formula><mml:math id="M345" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> period takes a maximum near <inline-formula><mml:math id="M346" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>. This demonstrates that the 95 <inline-formula><mml:math id="M347" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles in the G24-3 model are generated via nonlinear resonance.</p>
      <p id="d2e5634">We note that the closeness between the intrinsic timescale and the 95 <inline-formula><mml:math id="M348" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity period not only ensures the <inline-formula><mml:math id="M349" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M350" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> dominant period of ice age cycles but also enhances the temporal consistency between the simulations and the proxy data, as shown by Pearson's correlation coefficients for varying parameters <inline-formula><mml:math id="M351" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula> and <inline-formula><mml:math id="M352" display="inline"><mml:mi>A</mml:mi></mml:math></inline-formula> in Fig. S5 in the Supplement.</p>
</sec>
</sec>
<sec id="Ch1.S4">
  <label>4</label><title>Discussion</title>
      <p id="d2e5686">Our sensitivity experiments show that the models' responses can lock into individual or combined astronomical frequencies, depending on their intrinsic timescales (Fig. <xref ref-type="fig" rid="F5"/>). The locking frequency can also depend on the amplitude of the astronomical forcing (Fig <xref ref-type="fig" rid="F5"/>b and c). However, under realistic forcing amplitudes, models tend to produce <inline-formula><mml:math id="M353" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M354" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles when their intrinsic timescales are close to 100 <inline-formula><mml:math id="M355" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. This reflects a general property of synchronization and nonlinear resonance, observed across many ice age models (Table <xref ref-type="table" rid="T1"/>).</p>

<table-wrap id="T1" specific-use="star"><label>Table 1</label><caption><p id="d2e5725">Intrinsic timescales of models simulating <inline-formula><mml:math id="M356" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M357" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> glacial cycles. <inline-formula><mml:math id="M358" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> is the period of a self-sustained oscillation, <inline-formula><mml:math id="M359" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>nat</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> is the period corresponding to the natural frequency, and <inline-formula><mml:math id="M360" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> is the timescale for forming a threshold-triggered cycle. The asterisks (*) indicate the models explored in this study.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="3">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="left"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Model</oasis:entry>
         <oasis:entry colname="col2">Timescale <inline-formula><mml:math id="M361" display="inline"><mml:mrow class="unit"><mml:mo>(</mml:mo><mml:mi mathvariant="normal">kyr</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">Type of dynamics</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">
                  <xref ref-type="bibr" rid="bib1.bibx57" id="text.58"/>
                </oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M362" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub><mml:mo>≈</mml:mo><mml:mn mathvariant="normal">80</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">synchronization of a sustained oscillator</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">
                  <xref ref-type="bibr" rid="bib1.bibx73" id="text.59"/>
                </oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M363" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">98</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">synchronization of a sustained oscillator</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">
                  <xref ref-type="bibr" rid="bib1.bibx27" id="text.60"/>
                </oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M364" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub><mml:mo>≈</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">synchronization of a sustained oscillator</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">
                  <xref ref-type="bibr" rid="bib1.bibx17" id="text.61"/>
                </oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M365" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">103</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">synchronization of a sustained oscillator</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">
                  <xref ref-type="bibr" rid="bib1.bibx53" id="text.62"/>
                </oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M366" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">119</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">synchronization of a sustained oscillator</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">
                  <xref ref-type="bibr" rid="bib1.bibx3" id="text.63"/>
                </oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M367" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub><mml:mo>≃</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">synchronization of a sustained oscillator</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><xref ref-type="bibr" rid="bib1.bibx25" id="text.64"/> model 1</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M368" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">101.2</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">synchronization of a sustained oscillator</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">*SO model (present study)</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M369" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>SO</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">91.7</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">synchronization of a sustained oscillator</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">*<xref ref-type="bibr" rid="bib1.bibx78" id="text.65"/></oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M370" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>nat</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">95</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">nonlinear resonance in a damped oscillatory system</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">
                  <xref ref-type="bibr" rid="bib1.bibx6" id="text.66"/>
                </oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M371" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>≈</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">stochastic resonance in a bistable system</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">
                  <xref ref-type="bibr" rid="bib1.bibx60" id="text.67"/>
                </oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M372" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>≈</mml:mo><mml:mn mathvariant="normal">148</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">regime transitions at threshold crossings</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">*<xref ref-type="bibr" rid="bib1.bibx25" id="text.68"/> model 3</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M373" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>cyc</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">76.5</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">regime transitions at threshold crossings</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d2e6152">For example, <xref ref-type="bibr" rid="bib1.bibx60" id="text.69"/> simulate glacial cycles using a threshold-based regime-switching model, where the inherent time until the ice increases from <inline-formula><mml:math id="M374" display="inline"><mml:mrow><mml:msub><mml:mi>v</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">4.5</mml:mn></mml:mrow></mml:math></inline-formula> to <inline-formula><mml:math id="M375" display="inline"><mml:mrow><mml:msub><mml:mi>v</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">123</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M376" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula> is <inline-formula><mml:math id="M377" display="inline"><mml:mrow><mml:mo>(</mml:mo><mml:msub><mml:mi>v</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>v</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>)</mml:mo><mml:mo>/</mml:mo><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi>g</mml:mi></mml:msub><mml:mo>≃</mml:mo><mml:mn mathvariant="normal">126</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M378" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, and the timescale for deglaciation is <inline-formula><mml:math id="M379" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">d</mml:mi></mml:msub><mml:mi>ln⁡</mml:mi><mml:mfenced close="|" open="|"><mml:mrow><mml:mo>(</mml:mo><mml:msub><mml:mi>v</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi mathvariant="normal">d</mml:mi></mml:msub><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">d</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mo>/</mml:mo><mml:mo>(</mml:mo><mml:msub><mml:mi>v</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi mathvariant="normal">d</mml:mi></mml:msub><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi mathvariant="normal">d</mml:mi></mml:msub><mml:mo>)</mml:mo></mml:mrow></mml:mfenced><mml:mo>≃</mml:mo><mml:mn mathvariant="normal">22</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M380" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. Thus, the timescale for forming a cycle is 148 <inline-formula><mml:math id="M381" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. Although longer than 100 <inline-formula><mml:math id="M382" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, it is still closer to 100 <inline-formula><mml:math id="M383" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> than 20, 41, or 400 <inline-formula><mml:math id="M384" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. <xref ref-type="bibr" rid="bib1.bibx6" id="text.70"/> consider noise-induced transitions between wells under weak 100 <inline-formula><mml:math id="M385" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodic forcing. In the model, the signal-to-noise ratio is maximal at a certain noise intensity, the so-called <italic>stochastic resonance</italic>. This occurs when the average waiting time between two noise-induced transitions between the two wells (the inverse of the Kramers rate) is half the forcing period, i.e., <inline-formula><mml:math id="M386" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">50</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M387" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx5" id="paren.71"/>. Therefore, the intrinsic timescale of a cycle is <inline-formula><mml:math id="M388" display="inline"><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>×</mml:mo><mml:mn mathvariant="normal">50</mml:mn><mml:mo>=</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M389" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. The stochastic resonance theory is one of earliest examples treating the <inline-formula><mml:math id="M390" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M391" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> problem as a matching problem between the Earth's intrinsic timescale and external astronomical timescale. This theory has since been extended to align with Milankovitch theory <xref ref-type="bibr" rid="bib1.bibx48 bib1.bibx22" id="paren.72"/>. On the other hand, the piecewise linear model by <xref ref-type="bibr" rid="bib1.bibx35" id="text.73"/> is the example of a model with no 100 <inline-formula><mml:math id="M392" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> scale intrinsic timescale, and it fails to simulate the dominant <inline-formula><mml:math id="M393" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M394" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity. Its intrinsic timescales are 42.5 <inline-formula><mml:math id="M395" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> for glaciation and 10.6 <inline-formula><mml:math id="M396" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> for deglaciation.</p>
      <p id="d2e6472">The models of <inline-formula><mml:math id="M397" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M398" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles mentioned above are simple conceptual models, but some studies in the literature offer insights on how our timescale-matching hypothesis may hold in more complex models. First, an early study by <xref ref-type="bibr" rid="bib1.bibx57" id="text.74"/> demonstrated that an ice-sheet–bedrock system could exhibit 100 <inline-formula><mml:math id="M399" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> scale self-sustained oscillations, especially due to strong feedbacks involving basal melting and sliding of the ice sheets. This model serves as an example in which 100 <inline-formula><mml:math id="M400" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> scale intrinsic oscillations are relevant for producing 100 <inline-formula><mml:math id="M401" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles under insolation forcing, even though our understanding of ice-sheet and lithosphere physics has since been refined. Second, since the G24-3 model was, according to its author, inspired by experiments using the Earth system model of intermediate complexity, CLIMBER-2 model <xref ref-type="bibr" rid="bib1.bibx25" id="paren.75"/>, our results obtained from the G24-3 model can be relevant with complex climate systems including carbon cycles and dust–albedo interactions. <xref ref-type="bibr" rid="bib1.bibx54" id="text.76"/> showed that a version of the CLIMBER-2 model exhibits self-sustained oscillations with periods of several hundred thousand years, due to the glaciogenic dust feedback and carbon cycle feedbacks. Such long timescales are crucial for <inline-formula><mml:math id="M402" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M403" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> ice age cycles simulated in the CLIMBER-2 model under the forcing.</p>
      <p id="d2e6545">Although many models have intrinsic timescales around 100 <inline-formula><mml:math id="M404" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> (Table 1), not all have been evaluated from this perspective. The existence of an underlying 100 <inline-formula><mml:math id="M405" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> scale intrinsic timescale is hence our <italic>hypothesis</italic> based on the finite set of simple models surveyed here.</p>
      <p id="d2e6567">Different dynamical mechanisms add distinct nuance to the timescale-matching hypothesis. In the synchronization mechanism, the period of glacial cycles closely follows that of self-sustained oscillations, as suggested by the nearly vertical quasi-Arnold tongues (Fig. <xref ref-type="fig" rid="F5"/>a). In contrast, in the nonlinear resonance mechanism with damped oscillations, the natural period leading to the <inline-formula><mml:math id="M406" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M407" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles can deviate from 100 <inline-formula><mml:math id="M408" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, depending on the forcing amplitude, as suggested by the tilted 95 <inline-formula><mml:math id="M409" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> resonance region (Fig. <xref ref-type="fig" rid="F5"/>b). Thus, this mechanism does not require a precise match between the internal and external periods but rather a general alignment of their timescales. In the case of the bistable model (G24-3), the intrinsic timescale is not purely internal but includes a <inline-formula><mml:math id="M410" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">10</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M411" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> scale lag before favorable astronomical conditions are met. Although these mechanisms differ in their implications, the common factor for the emergence of the <inline-formula><mml:math id="M412" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M413" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles is the closeness of Earth's intrinsic timescale to the <inline-formula><mml:math id="M414" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M415" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity of the eccentricity cycles.</p>
      <p id="d2e6664">In this study, we distinguished between ice age models that exhibit synchronization and those that exhibit resonance. This distinction can be subtle in some cases. (i) In synchronization theory, the forcing is generally assumed to be small relative to the underlying self-oscillatory dynamics <xref ref-type="bibr" rid="bib1.bibx63" id="paren.77"/>. If the forcing is strong, it can significantly alter the oscillation amplitude, making it challenging to categorize the phenomenon strictly as either synchronization or resonance. (ii) <italic>Excitable systems</italic>, which are mono- or multistable in the absence of forcing, can produce repetitive oscillations when subject to small forcing or noise. If the frequency of such excited oscillations becomes locked to astronomical forcing, it resembles synchronization, though synchronization is typically reserved for systems with intrinsic self-sustained oscillations <xref ref-type="bibr" rid="bib1.bibx63" id="paren.78"/>. <xref ref-type="bibr" rid="bib1.bibx62" id="text.79"/> discusses the 100 <inline-formula><mml:math id="M416" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles from the perspective of a deterministic excitation paradigm and reaches a conclusion similar to ours: that an intrinsic timescale of <inline-formula><mml:math id="M417" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M418" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> should exist.</p>
      <p id="d2e6706">In <xref ref-type="bibr" rid="bib1.bibx78" id="text.80"/> as well as <xref ref-type="bibr" rid="bib1.bibx20" id="text.81"/>, the period doubling as well as the period tripling of the 41 <inline-formula><mml:math id="M419" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodic cycle is proposed as the scenario to give 100 <inline-formula><mml:math id="M420" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> scale glacial cycles (specifically 82 <inline-formula><mml:math id="M421" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> as well as 123 <inline-formula><mml:math id="M422" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles). This is not inconsistent with the present analysis of the VCV18 model. Indeed, if the VCV18 model is forced by the pure 41 <inline-formula><mml:math id="M423" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodic forcing, and if the forcing amplitude is increased, the period-doubling bifurcation is observed as shown in Fig. S6 in the Supplement. Comparing Fig. 5b with Fig. S6, the transition from the 41 <inline-formula><mml:math id="M424" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> regime to the 95 <inline-formula><mml:math id="M425" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> regime in Fig. 5b is considered an analog of a period-doubling bifurcation. The true period doubling is from 41 to 82 <inline-formula><mml:math id="M426" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>. However, the 95 <inline-formula><mml:math id="M427" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles are realized instead of the 82 <inline-formula><mml:math id="M428" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles because the climatic precession forcing, which is modulated by 95 <inline-formula><mml:math id="M429" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity cycles, is stronger than the obliquity forcing in the power of the summer solstice insolation at 65° N (Fig. <xref ref-type="fig" rid="F1"/>e).</p>
      <p id="d2e6807">Could the timescale-matching hypothesis be extended to the 41 <inline-formula><mml:math id="M430" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> dominant period observed before the Mid-Pleistocene Transition (MPT) <xref ref-type="bibr" rid="bib1.bibx8 bib1.bibx41" id="paren.82"/>? To address this question, it is important to recall that boreal summer insolation forcing contains only negligible <inline-formula><mml:math id="M431" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M432" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> power but a strong 41 <inline-formula><mml:math id="M433" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> component. If the 41 <inline-formula><mml:math id="M434" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> glacial cycles are simply linear responses, the intrinsic timescale may be less critical for their realizations. However, if they arise through synchronization or resonance, the intrinsic timescale becomes more relevant, as some models suggest.</p>
      <p id="d2e6857">In a study using the CLIMBER-2 model, <xref ref-type="bibr" rid="bib1.bibx54" id="text.83"/> found that 40 <inline-formula><mml:math id="M435" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> scale self-sustained oscillations underlie the 41 <inline-formula><mml:math id="M436" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> response prior to the MPT. In that study, the MPT is attributed to a gradual increase in the period of the self-sustained oscillations from <inline-formula><mml:math id="M437" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">40</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M438" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> to several hundred kyr. Timescale matching is key to the dominant period across the MPT. The G24-3 model exhibits a resonance scenario consistent with the timescale-matching hypothesis, showing shorter intrinsic timescales closer to 41 <inline-formula><mml:math id="M439" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> before the MPT and longer timescales near 76 <inline-formula><mml:math id="M440" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> after the MPT (Fig. S7 in the Supplement). However, the required proximity of the intrinsic timescale to 41 <inline-formula><mml:math id="M441" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> before the MPT depends on the model used. The VCV18 simulates the MPT-like transition if the parameters are changed in time so that the positive-to-negative feedback ratio is increased (Fig. S8a in the Supplement). Over the last 3 <inline-formula><mml:math id="M442" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula>, the natural period of damped oscillations increases from <inline-formula><mml:math id="M443" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">75</mml:mn></mml:mrow></mml:math></inline-formula> to 95 <inline-formula><mml:math id="M444" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, which is calculated from the complex eigenvalue of the Jacobian matrix at the stable state (Fig. S8 in the Supplement). Although the natural period before the MPT (75–80 <inline-formula><mml:math id="M445" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>) is still longer than the observed 41 <inline-formula><mml:math id="M446" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula>, this subtle shift is sufficient to produce a 41 <inline-formula><mml:math id="M447" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity in the VCV18 model. This behavior is already indicated by the 41 <inline-formula><mml:math id="M448" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> region adjacent to the 95 <inline-formula><mml:math id="M449" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> resonance region in Fig. <xref ref-type="fig" rid="F5"/>b. While models such as CLIMBER-2, G24-3, and VCV18 suggest that long-term parameter changes can shift the intrinsic timescale across the MPT, other models reproduce the MPT-like shift in dominant periodicity without requiring explicit parameter changes <xref ref-type="bibr" rid="bib1.bibx36 bib1.bibx33 bib1.bibx79" id="paren.84"/>. Investigating the relationship between the intrinsic timescale and the 41 <inline-formula><mml:math id="M450" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> response using the present approach requires comparing more models that accurately simulate the records through the MPT. We thus postpone this research to future work.</p>
</sec>
<sec id="Ch1.S5" sec-type="conclusions">
  <label>5</label><title>Summary</title>
      <p id="d2e7012">Determining origin of the <inline-formula><mml:math id="M451" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M452" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity in the Late Pleistocene glacial cycles has been an enduring problem in paleoclimate studies. We investigated three simple models of ice age cycles, which produce <inline-formula><mml:math id="M453" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M454" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles through distinct mechanisms: synchronization of self-sustained oscillations and nonlinear resonance in mono- or multi-stable systems. Although the astronomical forcing possesses only negligible power in the 100 <inline-formula><mml:math id="M455" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> band, these models exhibit <inline-formula><mml:math id="M456" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M457" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> ice age cycles as a response to the amplitude modulation of climatic precession cycles. This is physically equivalent to a response to the <inline-formula><mml:math id="M458" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M459" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> eccentricity cycles that modulate climatic precession. Through sensitivity experiments varying the intrinsic timescale of each model, we have revealed that the key factor for the emergence of the <inline-formula><mml:math id="M460" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M461" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> cycles is the closeness of the Earth's intrinsic timescale to the <inline-formula><mml:math id="M462" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M463" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> periodicity of the eccentricity cycles, regardless of the specific dynamical mechanism. In other words, the <inline-formula><mml:math id="M464" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">100</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M465" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kyr</mml:mi></mml:mrow></mml:math></inline-formula> period in the astronomical forcing is “selected” because it is close to the intrinsic timescale of the climate system. Note that this is a hypothesis derived from a finite set of models, mostly simple ones. Investigating the intrinsic timescales of more complex models is challenging. If adjusting the timescale of a model proves difficult, artificially varying the astronomical frequencies and observing the response could be a useful approach for evaluating the validity of the timescale-matching hypothesis in complex models.</p>
</sec>

      
      </body>
    <back><app-group>

<app id="App1.Ch1.S1">
  <label>Appendix A</label><title>Power spectral density method</title>
      <p id="d2e7162">The power spectral density (PSD) <inline-formula><mml:math id="M466" display="inline"><mml:mrow><mml:mi>S</mml:mi><mml:mo>(</mml:mo><mml:mi>f</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> of a time series is estimated using the periodogram <xref ref-type="bibr" rid="bib1.bibx11" id="paren.85"/>, which is computed with the <inline-formula><mml:math id="M467" display="inline"><mml:mi>R</mml:mi></mml:math></inline-formula> function <monospace>spec.pgram</monospace> <xref ref-type="bibr" rid="bib1.bibx64" id="paren.86"/>. By default, this function applies a split cosine bell taper to 10 <inline-formula><mml:math id="M468" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">%</mml:mi></mml:mrow></mml:math></inline-formula> of the data at both the beginning and the end of the time series to minimize discontinuity effects between the start and end of the series. To increase the number of frequency bins in the periodogram, zeros are added to the end of the series to extend its length by a factor of 10 (i.e., <monospace>pad=9</monospace> in the <monospace>spec.pgram</monospace> option). Zero padding does not fundamentally affect the PSD of the signal, but the frequency corresponding to a PSD peak is estimated with a higher resolution.</p>
</app>

<app id="App1.Ch1.S2">
  <label>Appendix B</label><title>van der Pol–Duffing–Hill equation</title>
      <p id="d2e7218">We assume that the glacial cycles can be represented by a forced van der Pol–Duffing–Hill equation:

              <disp-formula specific-use="align" content-type="numbered"><mml:math id="M469" display="block"><mml:mtable displaystyle="true"><mml:mtr><mml:mtd><mml:mstyle displaystyle="true" class="stylechange"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">¨</mml:mo></mml:mover><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>-</mml:mo><mml:mi mathvariant="italic">κ</mml:mi><mml:mo>)</mml:mo><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>+</mml:mo><mml:mi mathvariant="italic">α</mml:mi><mml:mi>x</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">β</mml:mi><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mo>+</mml:mo><mml:mi mathvariant="italic">θ</mml:mi></mml:mrow></mml:mtd></mml:mtr><mml:mlabeledtr id="App1.Ch1.S2.E3"><mml:mtd><mml:mtext>B1</mml:mtext></mml:mtd><mml:mtd><mml:mstyle displaystyle="true" class="stylechange"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">ν</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi mathvariant="italic">η</mml:mi><mml:msup><mml:mi>I</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mlabeledtr></mml:mtable></mml:math></disp-formula>

        where the parameters are denoted by Greek letters, and <inline-formula><mml:math id="M470" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is an insolation anomaly defined below. Under the restriction to the second-order differential equation, Eq. (<xref ref-type="disp-formula" rid="App1.Ch1.S2.E3"/>) is quite comprehensive from the viewpoint of dynamical systems. First, it contains the van der Pol equation <inline-formula><mml:math id="M471" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">¨</mml:mo></mml:mover><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>-</mml:mo><mml:mi mathvariant="italic">κ</mml:mi><mml:mo>)</mml:mo><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>+</mml:mo><mml:mi mathvariant="italic">α</mml:mi><mml:mi>x</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M472" display="inline"><mml:mi mathvariant="italic">κ</mml:mi></mml:math></inline-formula>, <inline-formula><mml:math id="M473" display="inline"><mml:mi mathvariant="italic">ν</mml:mi></mml:math></inline-formula>, and <inline-formula><mml:math id="M474" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> are typically positive <xref ref-type="bibr" rid="bib1.bibx77 bib1.bibx75" id="paren.87"/>. The van der Pol equation is well studied as a generic system showing self-sustained oscillations. Crucifix's group <xref ref-type="bibr" rid="bib1.bibx17 bib1.bibx18 bib1.bibx21 bib1.bibx52" id="paren.88"/> and others <xref ref-type="bibr" rid="bib1.bibx51 bib1.bibx4" id="paren.89"/> have used the forced van der Pol equation as a mathematical model for investigating ice age dynamics since it can roughly fit the Late Pleistocene glacial cycles.</p>
      <p id="d2e7424">Second, Eq. (<xref ref-type="disp-formula" rid="App1.Ch1.S2.E3"/>) contains the Hill equation <inline-formula><mml:math id="M475" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">¨</mml:mo></mml:mover><mml:mo>+</mml:mo><mml:mo>[</mml:mo><mml:mi mathvariant="italic">α</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>]</mml:mo><mml:mi>x</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula> if <inline-formula><mml:math id="M476" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is periodic in time <xref ref-type="bibr" rid="bib1.bibx46" id="paren.90"/>. Furthermore, if <inline-formula><mml:math id="M477" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is a simple harmonic, the Hill equation is called the Mathieu equation <inline-formula><mml:math id="M478" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">¨</mml:mo></mml:mover><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">α</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>cos⁡</mml:mi><mml:mn mathvariant="normal">2</mml:mn><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mi>x</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>. The latter is invoked to explain the rhythm of ice age cycles by <xref ref-type="bibr" rid="bib1.bibx67" id="text.91"/> from the viewpoint of frequency modulation.</p>
      <p id="d2e7538">Third, for <inline-formula><mml:math id="M479" display="inline"><mml:mrow><mml:mi mathvariant="italic">κ</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula>, Eq. (<xref ref-type="disp-formula" rid="App1.Ch1.S2.E3"/>) contains the Duffing equation <inline-formula><mml:math id="M480" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">¨</mml:mo></mml:mover><mml:mo>-</mml:mo><mml:mi mathvariant="italic">κ</mml:mi><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>+</mml:mo><mml:mi mathvariant="italic">α</mml:mi><mml:mi>x</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">β</mml:mi><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">ν</mml:mi><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> if <inline-formula><mml:math id="M481" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is a sinusoid. It is a paradigmatic system of nonlinear resonance as well as chaos <xref ref-type="bibr" rid="bib1.bibx24 bib1.bibx75" id="paren.92"/>. The Duffing equation exhibits forced oscillations but not self-sustained oscillations. Dropping out the additive forcing and the nonlinear damping term, Eq. (<xref ref-type="disp-formula" rid="App1.Ch1.S2.E3"/>) reduces to the model by <xref ref-type="bibr" rid="bib1.bibx20" id="text.93"/>: <inline-formula><mml:math id="M482" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">¨</mml:mo></mml:mover><mml:mo>+</mml:mo><mml:mi>a</mml:mi><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>-</mml:mo><mml:mi>b</mml:mi><mml:mi>x</mml:mi><mml:mo>+</mml:mo><mml:mi>b</mml:mi><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mo>+</mml:mo><mml:mi>b</mml:mi><mml:mi>c</mml:mi><mml:mo>-</mml:mo><mml:mi>b</mml:mi><mml:mi>x</mml:mi><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>,</mml:mo></mml:mrow></mml:math></inline-formula> where <inline-formula><mml:math id="M483" display="inline"><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover></mml:math></inline-formula> is the global temperature anomaly; <inline-formula><mml:math id="M484" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> represents a climatic memory effect; and <inline-formula><mml:math id="M485" display="inline"><mml:mi>a</mml:mi></mml:math></inline-formula>, <inline-formula><mml:math id="M486" display="inline"><mml:mi>b</mml:mi></mml:math></inline-formula>, and <inline-formula><mml:math id="M487" display="inline"><mml:mi>c</mml:mi></mml:math></inline-formula> are parameters (different symbols are used in the original reference).  Their model is essentially the Duffing–Hill equation since the damping term is linear. A modified version of their model can fit the proxy record well <xref ref-type="bibr" rid="bib1.bibx69" id="paren.94"/>.</p>
      <p id="d2e7733">A way to link Eq. (<xref ref-type="disp-formula" rid="App1.Ch1.S2.E3"/>) with a proxy variable of ice age cycles is to make a first-order system taking the so-called Liénard variable <inline-formula><mml:math id="M488" display="inline"><mml:mrow><mml:mi>y</mml:mi><mml:mo>=</mml:mo><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>-</mml:mo><mml:mi mathvariant="italic">κ</mml:mi><mml:mi>x</mml:mi><mml:mo>+</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mi mathvariant="italic">μ</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:mfrac></mml:mstyle><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx37 bib1.bibx17" id="paren.95"/>, which yields Eqs. (<xref ref-type="disp-formula" rid="Ch1.E1"/>) and (<xref ref-type="disp-formula" rid="Ch1.E2"/>). <inline-formula><mml:math id="M489" display="inline"><mml:mrow><mml:mi>y</mml:mi><mml:mo>=</mml:mo><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mtext>O</mml:mtext><mml:mo>-</mml:mo><mml:mi mathvariant="italic">δ</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:math></inline-formula> links the variable <inline-formula><mml:math id="M490" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> with the modeled <inline-formula><mml:math id="M491" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M492" display="inline"><mml:mrow class="unit"><mml:mo>(</mml:mo><mml:mi mathvariant="normal">‰</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> with an offset <inline-formula><mml:math id="M493" display="inline"><mml:mrow><mml:mi mathvariant="italic">δ</mml:mi><mml:mo>+</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:math></inline-formula>. The variable <inline-formula><mml:math id="M494" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> is an unobserved climate variable. In association with insolation forcing <inline-formula><mml:math id="M495" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M496" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> determines whether the system is in a glaciation phase or in a deglaciation phase. The scaled summer solstice insolation anomaly <inline-formula><mml:math id="M497" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is defined as <inline-formula><mml:math id="M498" display="inline"><mml:mrow><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>≡</mml:mo><mml:mo>(</mml:mo><mml:msub><mml:mi>F</mml:mi><mml:mtext>65N</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">495.7</mml:mn><mml:mo>)</mml:mo><mml:mo>/</mml:mo><mml:mn mathvariant="normal">24</mml:mn></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M499" display="inline"><mml:mrow><mml:msub><mml:mi>F</mml:mi><mml:mtext>65N</mml:mtext></mml:msub><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is the summer solstice insolation [<inline-formula><mml:math id="M500" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">Wm</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>] at <inline-formula><mml:math id="M501" display="inline"><mml:mrow><mml:mn mathvariant="normal">65</mml:mn><mml:mi mathvariant="italic">°</mml:mi></mml:mrow></mml:math></inline-formula> N calculated with the solar constant of 1368 <inline-formula><mml:math id="M502" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">Wm</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> (Fig. 1c) <xref ref-type="bibr" rid="bib1.bibx40 bib1.bibx19" id="paren.96"/>. The nonlinear effect of the insolation, <inline-formula><mml:math id="M503" display="inline"><mml:mrow><mml:mi mathvariant="italic">η</mml:mi><mml:msup><mml:mi>I</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, is included to account for the lower sensitivity of the ice volume in the cold period <xref ref-type="bibr" rid="bib1.bibx59" id="paren.97"/>. The term <inline-formula><mml:math id="M504" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>x</mml:mi><mml:mi>I</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is a multiplicative forcing. Such a multiplicative term can appear, from physical point of view, in the energy balance via albedo effects, the ice-mass balance via temperature–precipitation feedback <xref ref-type="bibr" rid="bib1.bibx42" id="paren.98"/>, and the calcifier–alkalinity model <xref ref-type="bibr" rid="bib1.bibx58" id="paren.99"/>.</p>
      <p id="d2e8041">The parameters of the equations are calibrated so as to minimize the mean squared error over the last 1 <inline-formula><mml:math id="M505" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Myr</mml:mi></mml:mrow></mml:math></inline-formula>. The minimization is conducted with the Nelder–Mead method implemented in R function optim <xref ref-type="bibr" rid="bib1.bibx64" id="paren.100"/>. The resultant parameters are <inline-formula><mml:math id="M506" display="inline"><mml:mrow><mml:mi mathvariant="italic">κ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1.0536394044</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M507" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">2.9662458029</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M508" display="inline"><mml:mrow><mml:mi mathvariant="italic">α</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.0356079021</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M509" display="inline"><mml:mrow><mml:mi mathvariant="italic">β</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.0001000922</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M510" display="inline"><mml:mrow><mml:mi mathvariant="italic">θ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.0180996836</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M511" display="inline"><mml:mrow><mml:mi mathvariant="italic">ν</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.0514402004</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M512" display="inline"><mml:mrow><mml:mi mathvariant="italic">ρ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.0189082535</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M513" display="inline"><mml:mrow><mml:mi mathvariant="italic">η</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.0049923333</mml:mn></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M514" display="inline"><mml:mrow><mml:mi mathvariant="italic">δ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.1801349684</mml:mn></mml:mrow></mml:math></inline-formula>.</p>
</app>
  </app-group><notes notes-type="codedataavailability"><title>Code and data availability</title>

      <p id="d2e8168">The R package Palinsol is available from CRAN. Additional R codes used in this study are available at <uri>https://github.com/takahito321/Codes-for-ESD-paper-on-100-kyr-ice-age-cycles.git</uri> (last access: 24 September 2025) (<ext-link xlink:href="https://doi.org/10.5281/zenodo.17168388" ext-link-type="DOI">10.5281/zenodo.17168388</ext-link>, <xref ref-type="bibr" rid="bib1.bibx50" id="altparen.101"/>). The tuned and untuned LR04 benthic stack records are available from <uri>https://lorraine-lisiecki.com/stack.html</uri> (last access:  2 December 2024). The <xref ref-type="bibr" rid="bib1.bibx31" id="text.102"/>  composite <inline-formula><mml:math id="M515" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> record on the depth-derived age model is available from <uri>https://www.ncei.noaa.gov/pub/data/paleo/contributions_by_author/huybers2006/huybers2006.txt</uri> (last access: 2 December 2024).</p>
  </notes><app-group>
        <supplementary-material position="anchor"><p id="d2e8203">The supplement related to this article is available online at <inline-supplementary-material xlink:href="https://doi.org/10.5194/esd-16-1569-2025-supplement" xlink:title="pdf">https://doi.org/10.5194/esd-16-1569-2025-supplement</inline-supplementary-material>.</p></supplementary-material>
        </app-group><notes notes-type="authorcontribution"><title>Author contributions</title>

      <p id="d2e8212">MC and PD provided the original research plan, which was merged with another plan by TM and NB. PD and TM extended the van der Pol-type oscillator model introduced by MC <xref ref-type="bibr" rid="bib1.bibx17" id="paren.103"/>. TM performed the simulation and numerical analysis, with substantial contributions from the others. All authors contributed to discussing the results and analysis throughout the research. The manuscript was written by all authors, with TM preparing the first draft.</p>
  </notes><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d2e8221">At least one of the (co-)authors is a member of the editorial board of <italic>Earth System Dynamics</italic>. The peer-review process was guided by an independent editor, and the authors also have no other competing interests to declare.</p>
  </notes><notes notes-type="disclaimer"><title>Disclaimer</title>

      <p id="d2e8230">Publisher's note: Copernicus Publications remains neutral with regard to jurisdictional claims made in the text, published maps, institutional affiliations, or any other geographical representation in this paper. While Copernicus Publications makes every effort to include appropriate place names, the final responsibility lies with the authors.</p>
  </notes><ack><title>Acknowledgements</title><p id="d2e8236">The authors thank Mikhail Verbitsky for his valuable comments during interactive discussions. Takahito Mitsui thanks Matteo Willeit for valuable discussions and Keita Tokuda for his kind support.</p></ack><notes notes-type="financialsupport"><title>Financial support</title>

      <p id="d2e8241">Takahito Mitsui and Niklas Boers have received funding from the Volkswagen Foundation. This is ClimTip contribution no. 27; the ClimTip project has received funding from the European Union's Horizon Europe research and innovation program (grant no. 101137601). Niklas Boers has received further funding from the European Union's Horizon 2020 research and innovation program under the Marie Skłodowska-Curie grant agreement no. 956170, as well as the Federal Ministry of Education and Research (grant no. 01LS3001A). This work was supported by JSPS KAKENHI (grant number 25K07942).</p>
  </notes><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d2e8247">This paper was edited by Claudia Pasquero and reviewed by Holger Kantz and one anonymous referee.</p>
  </notes><ref-list>
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