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  <front>
    <journal-meta><journal-id journal-id-type="publisher">ESD</journal-id><journal-title-group>
    <journal-title>Earth System Dynamics</journal-title>
    <abbrev-journal-title abbrev-type="publisher">ESD</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Earth Syst. Dynam.</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">2190-4987</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/esd-12-671-2021</article-id><title-group><article-title>Earth system economics: a biophysical approach <?xmltex \hack{\break}?> to the human component of the Earth system</article-title><alt-title>Earth system economics</alt-title>
      </title-group><?xmltex \runningtitle{Earth system economics}?><?xmltex \runningauthor{E.~D.~Galbraith}?>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1 aff2 aff3">
          <name><surname>Galbraith</surname><given-names>Eric D.</given-names></name>
          <email>eric.galbraith@mcgill.ca</email>
        <ext-link>https://orcid.org/0000-0003-4476-4232</ext-link></contrib>
        <aff id="aff1"><label>1</label><institution>Department of Earth and Planetary Science, McGill University, Montréal,  Canada</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Institut de Ciència i Tecnologia Ambientals (ICTA-UAB), Universitat Autònoma de Barcelona, <?xmltex \hack{\break}?> Barcelona, Spain</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>ICREA, Barcelona, Spain</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Eric D. Galbraith (eric.galbraith@mcgill.ca)</corresp></author-notes><pub-date><day>27</day><month>May</month><year>2021</year></pub-date>
      
      <volume>12</volume>
      <issue>2</issue>
      <fpage>671</fpage><lpage>687</lpage>
      <history>
        <date date-type="received"><day>27</day><month>August</month><year>2020</year></date>
           <date date-type="accepted"><day>8</day><month>April</month><year>2021</year></date>
           <date date-type="rev-recd"><day>5</day><month>April</month><year>2021</year></date>
           <date date-type="rev-request"><day>4</day><month>September</month><year>2020</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2021 Eric D. Galbraith</copyright-statement>
        <copyright-year>2021</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021.html">This article is available from https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021.html</self-uri><self-uri xlink:href="https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021.pdf">The full text article is available as a PDF file from https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021.pdf</self-uri>
      <abstract><title>Abstract</title>
    <p id="d1e98">The study of humans has largely been carried out in isolation from the study of the non-human Earth system. This isolation has encouraged the development of incompatible philosophical, aspirational, and methodological approaches that have proven very difficult to integrate with those used for the non-human remainder of the Earth system. Here, an approach is laid out for the scientific study of the global human system that is intended to facilitate seamless integration with non-human processes by striving for a consistent physical basis, for which the name Earth system economics is proposed. The approach is typified by a foundation on state variables, central among which is the allocation of time amongst activities by human populations, and an orientation towards considering human experience. A framework is elaborated which parses the Earth system into six classes of state variables, including a neural structure class that underpins many essential features of humanity.  A working example of the framework is then illustrated with a simple numerical model, considering a global population that is engaged in one of two waking activities: provisioning food or doing something else. The two activities are differentiated by their motivational factors, outcomes on state variables, and associated subjective experience. While the illustrative model is a gross simplification of reality, the results suggest how neural characteristics and subjective experience can emerge from model dynamics.  The approach is intended to provide a flexible and widely applicable strategy for understanding the human–Earth system, appropriate for physically based assessments of the past and present, as well as contributing to long-term model projections that are naturally oriented towards improving human well-being.</p>
  </abstract>
    </article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction</title>
      <p id="d1e110">Over the past 4 decades, Earth system science has developed a rich
understanding of interactions between the myriad physical, chemical, and
biological components of our planet <xref ref-type="bibr" rid="bib1.bibx57" id="paren.1"/>. By
considering the Earth as a single system, which is itself comprised of a
hierarchy of mechanistically interacting subsystems, Earth system science has
facilitated the challenge of thinking across vast scales of space and time and contextualized global change within the long-term evolution of life
<xref ref-type="bibr" rid="bib1.bibx39" id="paren.2"/>. In its quest to understand planetary functioning, this new
science has succeeded in crossing many disciplinary boundaries, developing
entirely new approaches – such as global carbon cycle science
<xref ref-type="bibr" rid="bib1.bibx23" id="paren.3"/> – and, in return, has brought fresh thinking to
the previously isolated disciplines from which it was born.</p>
      <p id="d1e122">Yet, despite being motivated by the human impacts on the planet, Earth system
science has done relatively little to directly incorporate humans themselves
<xref ref-type="bibr" rid="bib1.bibx45 bib1.bibx21 bib1.bibx14" id="paren.4"/>. For example, although
the seminal textbook by <xref ref-type="bibr" rid="bib1.bibx37" id="text.5"/> discusses human impacts on the planet
at length, there is no mention of human demographics, societal dynamics, or
well-being. Instead, the impacts of the human system<?pagebreak page672?> are viewed as external
forcings on the non-human Earth. This exclusion is particularly clear when
considering Earth system models (ESMs), the numerical flagships of Earth
system science. ESMs encapsulate the current understanding of the planet by
representing the component systems in a simplified fashion, integrated within
a seamless framework and discretized on a global grid. Because all component
systems co-exist within the same spatial framework and because they are based
on common foundations of biology, chemistry, and physics, the means of exchange
between the component systems are obvious, so that they can be integrated as a
whole to provide a synoptic global view. But ESMs do not include the global
human system within the same common foundations, and, as a result, the synoptic
perspective of Earth system science typically fails to include its most
rapidly changing and disruptive component <xref ref-type="bibr" rid="bib1.bibx44" id="paren.6"/>. This is not to say
there are no efforts in this direction; for example macroeconomic models are
frequently run in parallel with ESMs while exchanging information on
greenhouse gas emissions and increasingly sophisticated land use changes
<xref ref-type="bibr" rid="bib1.bibx49" id="paren.7"/>, an approach known as integrated assessment
modeling. But this approach is targeted primarily toward weighing 21st-century climate change impacts vs. greenhouse gas mitigation strategies
<xref ref-type="bibr" rid="bib1.bibx46 bib1.bibx64 bib1.bibx4" id="paren.8"/> rather
than to provide fundamental insight into the global human system. “Sectoral”
models often resolve geographically explicit interactions between humans and
specific Earth system components, such as the agricultural system
<xref ref-type="bibr" rid="bib1.bibx48" id="paren.9"/> or global marine fishery
<xref ref-type="bibr" rid="bib1.bibx27" id="paren.10"/>, but the fragmented sectoral approach does not
naturally build toward what <xref ref-type="bibr" rid="bib1.bibx65" id="text.11"/> call an
“integral” perspective on the human Earth.</p>
      <p id="d1e150">Why has there not been more effort devoted to the human component of Earth
system science? I suggest three reasons (though there are certainly
more). First, biologically identical humans have interacted with the Earth
system in very different ways when living under different social and
technological contexts. For example, a hunter–gatherer society has extremely
different per capita impacts on the Earth system than does a 21st-century
urbanized society. Thus, assuming a fixed set of functional characteristics
for our species – a strategy that works quite well for other organisms within
the Earth system – fails to address the most remarkable features of
humanity. Second, we care a lot about what humans think and how they feel,
which can make scientists hesitate to simplify features of humanity in the way
frequently done for other components of the Earth system. Third, there is a
vast cultural gulf between natural and social sciences that is very difficult
to bridge due to profoundly incompatible literatures. This gulf has left each
culture largely ignorant of the other, a problem that was identified decades
ago <xref ref-type="bibr" rid="bib1.bibx55" id="paren.12"/> and continues to persist.</p>
      <p id="d1e156">At the root of the natural–social science divide lies the difficulty of
linking the essential features of humanity – including knowledge systems,
social behaviour, and experience – to physical embodiments. This may reflect
the historical development of social sciences and humanities, originating as
they did when virtually all people believed in an eternal, disembodied soul
<xref ref-type="bibr" rid="bib1.bibx42" id="paren.13"/>. Thus, although many modern social scientists probably do
not subscribe to this belief, the underlying conceptual frameworks and
approaches remain aligned with its tacit implications, and many core features
of social science, such as values, beliefs, and norms, continue to depart from
non-physical starting points <xref ref-type="bibr" rid="bib1.bibx12" id="paren.14"/>. Differences in these
non-physical starting points have led, in turn, to a plethora of fields of
human study, among which there is little common ground, hampering
interdisciplinary progress.</p>
      <p id="d1e166">Yet, like all living organisms, humans are physical beings. The biological
reality of human bodies embeds us within ecosystems and links us to
biogeochemical cycles through our food production, material fluxes, and waste
flows <xref ref-type="bibr" rid="bib1.bibx32" id="paren.15"/>. The fact that each of us can be only in one place at
a time and engage in a limited number of activities per day places
fundamental physical constraints on our economies <xref ref-type="bibr" rid="bib1.bibx11" id="paren.16"/>. In
addition, advances in neuroscience now provide rich and compelling evidence
that everything that once appeared to be attributable to a disembodied soul is
actually formed “by the meat”, i.e. as emergent properties of our brains
<xref ref-type="bibr" rid="bib1.bibx15" id="paren.17"/>. The intricate network of synapses in each of our heads
determines what we think, how we feel, and who we are
<xref ref-type="bibr" rid="bib1.bibx38" id="paren.18"/>. These synapses are continually changing as we go through
our daily experiences, at rates that are biologically constrained
<xref ref-type="bibr" rid="bib1.bibx6" id="paren.19"/>. Thus, just as knowledge of the molecular processes
occurring within leaves can help to predict aspects of the global terrestrial
ecosystem <xref ref-type="bibr" rid="bib1.bibx58" id="paren.20"/>, there is good reason to hope that many aspects
of humanity, historically considered unquantifiable, can actually be better
understood by considering how they emerge from the physical constructs of
synaptic networks. Neuroscience still has much to learn about the functioning
of the brain, but it holds great promise as a common ground to help unify the
fragmented domains of social science <xref ref-type="bibr" rid="bib1.bibx13" id="paren.21"/>.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F1" specific-use="star"><?xmltex \currentcnt{1}?><?xmltex \def\figurename{Figure}?><label>Figure 1</label><caption><p id="d1e193">ESE provides a bridge between the Earth system science approach,
typified by Earth system models, and the diverse fields of human study.</p></caption>
        <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021-f01.png"/>

      </fig>

      <p id="d1e202">The lacklustre development of the human component of Earth system science is
also evident in its failure to enrich the scientific understanding of humans
themselves. This is in contrast to integrative Earth system approaches such as
ocean biogeochemistry, which has provided important insights into marine ecology
(e.g. <xref ref-type="bibr" rid="bib1.bibx24" id="altparen.22"/>).  Early work under the name of human
ecology made significant progress toward modeling hunter–gatherers through
their interactions with the environment (e.g. <xref ref-type="bibr" rid="bib1.bibx66" id="altparen.23"/>),
providing valuable insights for anthropology and sociology, but these works
were not widely seized upon. In contrast, there have been very few Earth-system-scale studies that ask fundamental questions about the physical
coupling of humans with the ecosystem <xref ref-type="bibr" rid="bib1.bibx45" id="paren.24"/> and even fewer
that explore the implications for the quality of human existence.<?pagebreak page673?> Yet human
well-being is of central importance to social science and policymakers and could either improve or deteriorate dramatically in future, depending on
societal choices <xref ref-type="bibr" rid="bib1.bibx10" id="paren.25"/>.</p>
      <p id="d1e217">Here a strategy is pursued to provide a seamless integration of human and
non-human parts by representing humans – like the rest of the Earth system – on a biophysical foundation. The strategy aims to facilitate new forms of
communication at the juncture of natural and social sciences, with the
aspiration of providing new insights for both human and non-human systems
(Fig. 1). The name Earth system economics (ESE) is proposed for the endeavour, though as discussed below, it differs from mainstream economics in a number of important ways. Section 2 gives a few examples of the types of problems that could be addressed with ESE. Section 3 provides an overview of the key guiding principles that motivate the ESE approach. Section 4 details a high-level conceptual framework for the global human system. Section 5 describes a  numerical model of the global human system, inspired by simple models of the global carbon cycle (e.g. <xref ref-type="bibr" rid="bib1.bibx52" id="altparen.26"/>), as an illustration of how the framework can be operationalized. Section 6 provides analysis and discussion of the model. Section 7 offers concluding comments.</p>
</sec>
<sec id="Ch1.S2">
  <label>2</label><title>Topics that could be readily addressed with Earth system economics</title>
      <p id="d1e231">As ESE pursues a new approach, it is difficult to foresee all applications that may arise from its development. Some of the more obvious applications might include the following:
<list list-type="order"><list-item>
      <p id="d1e236">gaining a mechanistic, bird's eye perspective on the global human system that allows for seamless analysis across scales, essentially capturing the human system as an integrated part of global material cycles and subject to physical constraints, with the ability to resolve high spatial resolution;</p></list-item><list-item>
      <p id="d1e240">suggesting new non-monetary metrics that capture key aspects of the global human system; despite long-standing calls for moving beyond gross domestic product (GDP) as the dominant measure of progress <xref ref-type="bibr" rid="bib1.bibx63 bib1.bibx17 bib1.bibx19" id="paren.27"/>, proposed alternatives are often converted to monetary units or subject to criticisms of arbitrariness. By focusing on physical quantities and their rates of change, ESE can offer an alternative set of objective, quantitative metrics to inform the human–Earth status;</p></list-item><list-item>
      <p id="d1e247">testing hypotheses in historical dynamics <xref ref-type="bibr" rid="bib1.bibx61" id="paren.28"/>; in the same way that Earth system models can be used to test hypotheses about past climate changes, ESE models could be applied to test hypotheses about past changes in the human system – for example, what emergent features are required to accurately hindcast the spatial progressions of key societal transitions in history, such as the Neolithic transitions or industrialization?</p></list-item><list-item>
      <p id="d1e254">the spatial and temporal dynamics of human interactions with ecosystems and consequences for biodiversity; tight coupling with physically based biodiversity models can provide new tools with which to test hypotheses regarding early mass extinctions or the controls on future threats to ecosystem stability in a spatially explicit context;</p></list-item><list-item>
      <p id="d1e258">mechanistic linkages between subjective well-being and the biophysical consequences of societal actions; how could human lived experience vary given different societal pathways and within physical constraints, including coupled Earth system impacts such as climate change and biodiversity loss?</p></list-item></list></p>
      <?pagebreak page674?><p id="d1e261"><?xmltex \hack{\newpage}?>Most of these complex problems have been addressed by other means, especially
at local scales, but all remain incompletely resolved. The ESE approach aims to provide a novel global, integrated view, while prompting new avenues for mechanistic insight. The approach may ultimately be more widely applicable than indicated by this short list.</p>
</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Guiding principles of Earth system economics</title>
      <p id="d1e273">In a nutshell, ESE aims to quantify physical aspects of the human–Earth system
(state variables), including how the physical state is dynamically altered by
human actions (time allocation) and consequences for the nature of human
experience. In this section, a few general principles are discussed, as
motivation for the framework which follows.</p>
<sec id="Ch1.S3.SS1">
  <label>3.1</label><title>Striving for physical foundations</title>
      <p id="d1e283">Foremost, ESE strives for a grounding in quantifiable, physical
terms. Physical variables exhibit persistence over time, and physical
processes impose firm limits on possible rates of change, leading to dynamic
predictability. Physical variables also lend themselves to strict definitions,
which can prevent double counting while simultaneously helping to ensure
inclusivity. Much of the predictive success of natural sciences lie in their
ultimate recourse to physical variables, which provide pathways to diverse
insights whether starting from biology, physics, or chemistry. For example, the
conservation of mass, momentum, and energy play essential roles in many
branches of Earth system science, from atmospheric circulation to ice sheet
motion and sea level rise. Ecosystem and biogeochemical models benefit from
the understandings of living processes as molecular interactions writ large.</p>
      <p id="d1e286">That said, the aim to ground the human system in physical quantities is not
trivial. For some things – population demographics, cars, infrastructure,
fossil fuel consumption – it is straightforward. But for many of the most
fascinating and important aspects – such as behavioural motivations and
subjective experience – the biophysical bases remain vaguely
described. Operationally, it will always be necessary to use coarse
approximations for these (and other) unresolved or poorly understood
processes, a common strategy in Earth system models, such as the
representation of cloud physics with empirical parameterizations. These
parameterizations are always unsatisfying, but the fact that they are
explicitly recognized as unsatisfactory and can ultimately be replaced by more
physically grounded mechanistic understandings identifies a direction for
progress. Resolutely abstract variables, on the other hand, resist connection
to complementary scientific insights and reinforce disciplinary silos. Thus,
the important thing is that strengthening the physical foundation is ever
present as a central goal of ESE: that long-term progress can be made by
improving the physical representation of all aspects of the human system
through improved observations and theoretical development.</p><?xmltex \hack{\newpage}?>
</sec>
<sec id="Ch1.S3.SS2">
  <label>3.2</label><title>Quantification of activities</title>
      <p id="d1e298">The diversity of human endeavours can be overwhelming and might appear to
defy a recourse to conserved quantities in the way that the motion of fluids
is linked to momentum and density through the Navier–Stokes equations. However, there is no question that the amount of time available to each human is a strictly conserved quantity. All humans engage in some form of activity for exactly 24 <inline-formula><mml:math id="M1" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">h</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. The activities in which a population is engaged determine its impact on the biophysical reality and also play a major role in determining the subjective experience of its individuals. Thus, activities are employed here as the central feature of ESE.</p>
      <p id="d1e318">There does not exist a universal system for classifying activities. Even the
activity of a reader of a scientific article can be described in many ways,
which may include reading, working, thinking, learning, sitting, or using a
screen or computer. The activity may be subjectively enjoyable or unpleasant,
depending on the quality of the text and disposition of the reader. The
optimal strategy to classify activities would involve as little subjective
interpretation as possible and be grounded as firmly as possible in physical
features, a possibility that could be further developed elsewhere. For the
moment, it is sufficient to consider this a difficult and
incompletely resolved problem.</p>
      <p id="d1e321">In the absence of a universal lexicon of activities, applicable to all humans
at all times, a lexicon must be constructed for a particular purpose. An
activity lexicon must identify, as unambiguously as possible, a set of
mutually exclusive activities that together include all possible activities
available to the population. Thus, the fractional distribution of time between
the activities must sum to exactly 1. For example, a simple two-activity
lexicon would be sleeping and not-sleeping. To be useful, the lexicon should
align activities with the outcomes that motivate them, by considering how they
modify state variables.</p>
</sec>
<sec id="Ch1.S3.SS3">
  <label>3.3</label><title>Subjective experience</title>
      <p id="d1e332">Humans live rich inner lives, and individuals can be either filled with joy or tormented by suffering, depending on what circumstances befall them. Improving the inner life experience of humans has pre-occupied much of society for generations and remains a central goal of global society, as exemplified by the UN Sustainable Development Goals: 11 of the 17 goals are
oriented towards improving the life experience of humans, while only six are oriented toward maintaining non-human aspects of the planet. Given that subjective experience appears to be a top priority for most of humanity, it is explicitly included as an essential component of the ESE approach.</p>
      <p id="d1e335">Despite its importance, the biophysical understanding of subjective experience
remains rudimentary (e.g. <xref ref-type="bibr" rid="bib1.bibx3" id="altparen.29"/>). It will take many years
of additional research before quantifications are available to assess human
experience that rival, for example, our ability to quantify the concentrations
of trace gases in the atmosphere. Nonetheless, the field of subjective
well-being has made great strides in providing large datasets on how people
themselves evaluate their life experiences <xref ref-type="bibr" rid="bib1.bibx20" id="paren.30"/>. These
can be considered along two axes:
<list list-type="order"><list-item>
      <p id="d1e346">Affect: this refers to the momentary emotions felt throughout the day, sometimes assessed by asking a subject whether they felt positive or negative emotions (e.g. laughed, cried, felt angry) over some preceding time interval <xref ref-type="bibr" rid="bib1.bibx18" id="paren.31"/> or by asking a candidate to rank the pleasantness <xref ref-type="bibr" rid="bib1.bibx28" id="paren.32"/> or unpleasantness <xref ref-type="bibr" rid="bib1.bibx36" id="paren.33"/> of different activities.</p></list-item><list-item>
      <p id="d1e359">Cognitive life evaluation and eudaemonia: these reflect time-integrated rather than momentary aspects of well-being. For cognitive life evaluations, the subject is asked to consider their life as a whole and evaluate their level of satisfaction with it, usually on a 10-point scale. The results are often correlated reasonably well with affect and can be predicted to some degree from material and non-material variables <xref ref-type="bibr" rid="bib1.bibx33 bib1.bibx10" id="paren.34"/>. The term eudaemonia refers to a fulfillment of purpose and is often oriented towards philosophical goals of what life ought to be rather than one that is desirable on purely hedonic terms <xref ref-type="bibr" rid="bib1.bibx51" id="paren.35"/>. Although a major concern of society on historical timescales, often addressed through religion, eudaemonia has been less studied in recent years, with less effort dedicated to developing quantitative indices.</p></list-item></list></p>
      <p id="d1e368">These axes of subjective well-being do not capture all that is important to
human experience, and the difficulty of comparing assessments between cultures
and languages cannot be taken lightly. But it appears likely that the
quantitative basis for constructing population-level assessments of life
experience will continue to improve as time progresses.</p>
</sec>
<?pagebreak page675?><sec id="Ch1.S3.SS4">
  <label>3.4</label><title>Drawing on all fields of human-related science</title>
      <p id="d1e380">Many disciplines study humans, including the core social sciences of
economics, anthropology, sociology, and psychology, as well as history,
medicine, law, business, and education. All of these disciplines can provide
useful insights on the global human system. For this reason, ESE aspires to
establish common ground that is compatible with aspects of all fields of human
study, by explicitly considering the physical foundations that underly them.</p>
      <p id="d1e383">So why use the term “economics”? In its modern use, this term has become
narrowly associated with the distribution of scarce resources, the production
and consumption of goods and services by firms and households, and monetary
exchanges. However, the origin of the word, from the Greek <italic>oikonomia</italic>,
referred to managing the home in a rational way in order to benefit its
occupants <xref ref-type="bibr" rid="bib1.bibx40" id="paren.36"/>. The root <italic>oiko</italic> is also the basis of
“ecology”, study of the home. The aim of the current proposal is to provide
an additional means for holistic, science-based perspectives to assist in
rational decision-making that can improve the management of the wealth of our
common home, the Earth system, for the benefit of its inhabitants. Hence, the
usage here is consistent with the original Greek term. Nonetheless, it should
be born in mind that ESE is only distantly related to mainstream economics.</p>
</sec>
<sec id="Ch1.S3.SS5">
  <label>3.5</label><title>Focus on population-level interactions</title>
      <p id="d1e403">ESE focuses on humans at the population level as the primary interactive
unit. Of course, human behaviours and experiences all actually happen at the
individual level. But just as the dynamics of a fluid can be usefully
described without resolving the motions of individual molecules within it,
population characteristics can be usefully described without resolving
individual interactions, and symmetry breaking can lead to fundamentally
different behaviour across scales <xref ref-type="bibr" rid="bib1.bibx5" id="paren.37"/>. What is more, these
population characteristics can show greater predictability when the emergent
result depends on well-behaved statistical distributions of individual
behaviour, as illustrated by the dynamics of human mobility
<xref ref-type="bibr" rid="bib1.bibx53 bib1.bibx1" id="paren.38"/>.</p>
      <p id="d1e412">Focusing on the population level does not mean that variability within the
population must be ignored. Variability can be incorporated as additional
information that describes the variability in a parameter, such as a
probability distribution function. For example, the distribution of wealth
within a population can often be approximated as a power law, for which only a
single parameter (the exponent) needs to be defined <xref ref-type="bibr" rid="bib1.bibx67" id="paren.39"/>.</p>
</sec>
<sec id="Ch1.S3.SS6">
  <label>3.6</label><title>Emphasis on the ultimate “what” and “why” of activities rather than the “how”</title>
      <p id="d1e426">A great deal of human study is oriented towards understanding how social
activities are coordinated and the means by which the cooperative activities
of many individuals can be optimized. The mechanisms by which this
coordination occurs underpin many fascinating aspects of culture, economics,
management, and law, but are not the target of enquiry here.</p>
      <p id="d1e429">Instead, ESE is characterized by a focus on the what and why of human
activities. Here, “what” refers to the final net outcome of an activity, or
complex of cooperative activities, in physical terms. The “why” refers to
the ultimate motivations for undertaking the activity, again in relation to
the final net outcome in the case of a complex of coordinated activities. For
example, the final outcome of creating farming tools, tilling soil, sowing
seeds, tending plants, and harvesting crops is to provide food (what). The
motivation for this<?pagebreak page676?> is a hunger-driven need for food (why). Thus, ESE aims
to circumvent the complexity of immediate, individual motivations for
component activities (such as whether the work is done for pay, which could
itself be motivated by material consumption, which itself could be motivated
by a desire to raise social standing) by considering the net outcome of any
set of activities as the relevant motivating factor.</p>
</sec>
<sec id="Ch1.S3.SS7">
  <label>3.7</label><title>Applicability to any point in time</title>
      <p id="d1e440">It could be easier to design a conceptual system exclusively for the
present day, with which we are intimately familiar, than one which works
equally well back to medieval times or the Late Pleistocene. Yet, if we aspire
to consider the distant future, many decades or centuries hence, this ability
must be a bare minimum requirement, since presumably the future could hold
many revolutionary changes that defy the imagination today. The ESE approach
strives to be applicable across the full temporal scope of human existence,
enabling hindcasts to test dynamical hypotheses against historical
observations as well as to explore hypothetical future projections.</p>
</sec>
<sec id="Ch1.S3.SS8">
  <label>3.8</label><title>Focus on emergent consequences of predictable aspects</title>
      <p id="d1e451">Most aspects of complex systems, including the human–Earth system, are
unpredictable. But within this sea of unpredictability lie islands of
predictability. For example, the chaotic processes that determine daily
weather can be approximated well enough to provide a very detailed forecast
over the next 12 h but are almost completely unpredictable on a
timescale of 1 month. Yet, on a coarser scale, seasonal and even decadal
climate forecasts are now reasonably good <xref ref-type="bibr" rid="bib1.bibx54" id="paren.40"/>.  Similarly,
societal dynamics include a vast variety of interacting, nonlinear processes
that are extremely challenging to predict but within which occur more
predictable aspects. Thus, ESE strives to identify the more robust, least
unpredictable aspects of the system, seeking insights on the emergent results
of their interactions. Societal, cultural, and economic characteristics of
populations are described through the simplifying lens of how they impact
physical variables and time allocation. The roles of the more unpredictable
aspects can then be assessed through the quantification of structural and
parameter uncertainty, the use of probability distributions, and the inclusion
of tipping points if they are identified through other means.</p>
</sec>
</sec>
<sec id="Ch1.S4">
  <label>4</label><title>Earth system economics conceptual framework</title>
      <p id="d1e466">Humans have an intellectual ability to foresee the future that is unparalleled
amongst other forms of life and an apparently infinite scope to modify their
biophysical surroundings. How could these features possibly be captured in a
numerical assessment? To paraphrase George Box, the answer is that it can be
done through countless ways, none of which is perfect but some of which can
be useful. And, as written in a discussion of Box's aphorism by
<xref ref-type="bibr" rid="bib1.bibx60" id="text.41"/>, “it may be necessary to create a model that takes a
totally different perspective in order to improve upon currently accepted
models.”</p>
      <p id="d1e472">Here, a new perspective on the human system is proposed that is consistent
with the ESE principles outlined in Sect. 3 and that forms an intuitive and
inclusive structure that aligns well with observational data. To be tractable
at the global scale, the framework definition is hierarchical, so that it can
be used at a high level of aggregation. The framework is inclusive,
encapsulating the entirety of the global human system, while aiming to
facilitate the representation of its mechanistic properties. At the same time,
the categories are conceptually straightforward to expand into disaggregated
detail, with as little ambiguity as possible, and spatial disaggregation
should be easy to apply. This proposed framework is intended as a
superstructure within which analyses or models could be developed through
further work.</p>
<sec id="Ch1.S4.SS1">
  <label>4.1</label><title>State variables</title>
      <p id="d1e482">The ESE framework is defined by state variables. Each state variable
represents some physical aspect of the human–Earth system, living or
non-living. Each variable could be measured and quantified over some spatial
and temporal domain (at least in theory, even if impractical or impossible
with current technology) and is subject to physical constraints.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F2"><?xmltex \currentcnt{2}?><?xmltex \def\figurename{Figure}?><label>Figure 2</label><caption><p id="d1e487">The superstructure of the conceptual framework. Each of the six shaded regions corresponds to a state variable class. Variables in the blue regions relate to the persistent structure and instantaneous activation of human neural systems, the orange region to human bodies, the brown region to human-created things, the grey to the distribution of time amongst activities, and the green to the remainder of the Earth system.</p></caption>
          <?xmltex \igopts{width=236.157874pt}?><graphic xlink:href="https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021-f02.png"/>

        </fig>

      <p id="d1e496">The highest-level grouping of six variable classes, proposed here, is
illustrated in Fig. 2 and elaborated below. The five variable classes in the
outer ring include everything on the surface of the Earth and can therefore
be thought of as a conceptual superstructure within which more detailed
subdivisions can be resolved. The final state variable class, time allocation,
is not physically embodied but is nonetheless subject to the limitation of
24 <inline-formula><mml:math id="M2" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">h</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> and is unambiguously defined for a population within a
given spatial–temporal domain.</p>
      <p id="d1e517"><italic>Soma</italic>. This refers to  the living ensemble of human bodies and their biophysical characteristics, including microbiota. The Soma determines the biogeochemical fluxes required to maintain the population, including food and water consumption as well as the production of heat and waste. It also includes properties reflecting the health status of the population (including symbiotic and pathogenic microbes) and physical fitness. Example state variables here could include the total population biomass (kg), an age-structured population description (number and age), or detailed information on body compositions (e.g. <inline-formula><mml:math id="M3" display="inline"><mml:mrow class="chem"><mml:mi mathvariant="normal">C</mml:mi><mml:mo>:</mml:mo><mml:mi mathvariant="normal">N</mml:mi></mml:mrow></mml:math></inline-formula> ratio, Fe content).</p>
      <?pagebreak page677?><p id="d1e534"><italic>Neural structure</italic>. It is because of the dynamical processes in our
brains that we are the dominant species on the planet. Our neurons encode
networks that are highly plastic, and this plasticity forms the foundation of
our ability to learn <xref ref-type="bibr" rid="bib1.bibx6" id="paren.42"/> as well as our responses to
stimuli <xref ref-type="bibr" rid="bib1.bibx41" id="paren.43"/>. The biophysical characteristics of our
brains lie at the foundation of core societal traits such as knowledge and
behaviours, as well as subjective experience <xref ref-type="bibr" rid="bib1.bibx41 bib1.bibx13" id="paren.44"/>. Thus, state variables describing the brains of humans within
a population can be used to represent these essential features. One type of
state variable could quantify aspects of associative links within the
population connectome (the ensemble of all synaptic connections in a
population, sensu <xref ref-type="bibr" rid="bib1.bibx56" id="altparen.45"/>), such as the number of
associations encoded by synapses. Other possibilities would be topological
descriptions of the neural structure, e.g. the degree of diversity of associations
within the population, the quality of the predictive capacity of associations,
or links with hormonal and emotional responses.  Although the processes
underlying the formation of new synapses and their selective destructions
remain incompletely understood, they are certain to happen at finite,
biologically constrained rates, placing limits on possible rates of learning
and modulating the persistence of behaviour, values, and emotional features
within a population. Conceptualizing neural structures as real, persistent
aspects of the Earth system prompts a novel perspective on the consequences of
human time allocation and points toward the underlying physical basis of how
systemic, population-level societal changes can occur.</p>
      <p id="d1e551"><?xmltex \hack{\newpage}?><italic>Neural activation</italic>. Existing neural structures are activated by
sensory stimulus and result in what we experience as thoughts, emotions, and
feelings. The sensory stimulus includes external factors such as the
landscape, music, food, mobile phone screens, and conversation, as well as
interoceptive body status such as hunger and thermal comfort, and in fact both
types of sources generally co-occur <xref ref-type="bibr" rid="bib1.bibx9" id="paren.46"/>. This class of
state variables represents manifestations of this neural activation. Because
the details of the activation itself remain difficult to observe, emergent
properties such as subjective well-being measures are most usable at present,
though observation technologies are rapidly improving. Neural activation is
also conceptually useful as the pathway by which neural structure is
modified.</p>
      <p id="d1e560"><italic>Things</italic>. Humans are clever, but it is not through individual
cleverness alone that we have become the dominant species on the planet
<xref ref-type="bibr" rid="bib1.bibx35" id="paren.47"/>. Rather, we leverage our ability to think by
creating entities with novel properties, constructed through shared knowledge
and social coordination, that then amplify our ability to modify the physical
environment. This includes the fabrication of tools, the construction of
buildings and infrastructure, the making of vehicles and airplanes, and the
writing of books and computer code. The Things class includes all of these and is defined as all non-living entities which are brought into existence as
a desired outcome of human activity. As such, the Things class does not
include livestock or genetically modified organisms, nor does it include
waste. Instead, these are regarded as modifications of the remainder of the
Earth system.</p>
      <p id="d1e568"><italic>Remainder of the Earth system</italic>. This includes all living organisms
other than humans (including agricultural plants and livestock), the
atmosphere, regolith, soil, rock, the ocean, and the cryosphere. These fall
within the traditional domain of Earth system science. Although the variables
within this class can all be affected by the human system, and many may be
very strongly modified (e.g. cows, grapefruit), they do not require human
activity in order to persist and/or are living organisms, thereby
differentiating them from Things.</p>
      <p id="d1e573"><italic>Time allocation</italic>. The allocation of time between activities is a
complex topic, which has been studied in many branches of social sciences (see
<xref ref-type="bibr" rid="bib1.bibx28" id="altparen.48"/>, for a useful overview). In a simple form, the
allocation of time can be regarded as the emergent outcome of competing
motivations, expressed at the population level. As discussed above, a
motivation is strictly defined as the reason to undertake an activity (the
why) that relates to the set of physical outcomes caused directly by the
activity (the what). The consequent population-level time allocation,
which emerges from the balance of competing motivations, causes changes in
state variables including subjective experience according to the context
(e.g. the presence of Things, neural structures, climate). The variables
in this class are simply the fraction of time (e.g. <inline-formula><mml:math id="M4" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">h</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>) devoted
to each activity by the population.</p>
</sec>
</sec>
<?pagebreak page678?><sec id="Ch1.S5">
  <label>5</label><title>Illustrative model</title>
      <p id="d1e608">Next, a simple model is presented that illustrates how the ESE framework could be operationalized in a global model. The model bears some similarity to simple ecological models that have previously aimed for direct coupling of
humans and ecosystems (e.g. <xref ref-type="bibr" rid="bib1.bibx34" id="altparen.49"/>) but using the ESE
conceptual basis. The model description follows the ODD (Overview, Design
concepts, Details) protocol for describing individual- and agent-based models
<xref ref-type="bibr" rid="bib1.bibx30" id="paren.50"/>, as updated by <xref ref-type="bibr" rid="bib1.bibx31" id="text.51"/>. Because the model
simulates changes in state variables according to differential equations,
rather than being agent-based, a number of the standard design concepts are
omitted from the description. Despite these omissions, the description is rather long, and the time-conscious reader may wish to skip most of this section.</p>
<sec id="Ch1.S5.SS1">
  <label>5.1</label><title>Purpose and patterns</title>
      <p id="d1e627">The purpose of the model is to illustrate how linkages can be simulated among
the six classes of state variables through dynamical interdependencies. The
model focuses on the interaction between a human population and an ecosystem
that provides food, with feedbacks mediated by state-dependent motivations
that re-allocate time between the activities of collecting food
(<italic>provisioning</italic>) and doing something else (<italic>other</italic>). The model generates time-varying patterns in population-level state variables, including neural structure and affect, and generalizes many societal features through their influences on motivating time allocation. This simple illustrative model is not intended to realistically capture any particular period of human history but to give an example of how the ESE conceptual framework could guide model construction.</p>
</sec>
<sec id="Ch1.S5.SS2">
  <label>5.2</label><title>Entities, state variables, and scales</title>
      <p id="d1e644">The model is not agent-based and therefore does not simulate interactions
amongst entities. Instead, the model considers dynamical changes in state
variables, starting from an initial state, according to ordinary differential
equations. This distinction is analogous to the distinction between what are
referred to as Eulerian and Lagrangian models in fluid dynamics
<xref ref-type="bibr" rid="bib1.bibx62" id="paren.52"/>. The approach taken here is equivalent to the Eulerian
method, modeling the human population as a field, characterized by its
state, rather than trying to resolve the motions of individual particles.</p>
      <p id="d1e650">The model includes state variables that fall within the six classes introduced
in Fig. 2, as listed in Table 1 and shown in Fig. 3. Each state variable has a
single scalar value at any point in time. The two time allocation variables
are <inline-formula><mml:math id="M5" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M6" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>. Each of these represents a
compound activity, comprised of many unresolved component activities.</p>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T1"><?xmltex \currentcnt{1}?><label>Table 1</label><caption><p id="d1e678">Model state variables.</p></caption><oasis:table frame="topbot"><?xmltex \begin{scaleboxenv}{.950}[.950]?><oasis:tgroup cols="3">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="left"/>
     <oasis:thead>
       <oasis:row>
         <oasis:entry colname="col1">Variable</oasis:entry>
         <oasis:entry colname="col2">Symbol</oasis:entry>
         <oasis:entry colname="col3">Value and</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">units</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">Human biomass</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M7" display="inline"><mml:mrow><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M8" display="inline"><mml:mrow class="unit"><mml:msub><mml:mi mathvariant="normal">kg</mml:mi><mml:mi mathvariant="normal">human</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Neural structure, provisioning-activated</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M9" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">no.</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Neural structure, other-activated</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M10" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M11" display="inline"><mml:mi mathvariant="italic">#</mml:mi></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Provisioning tools</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M12" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M13" display="inline"><mml:mrow class="unit"><mml:msub><mml:mi mathvariant="normal">kg</mml:mi><mml:mi mathvariant="normal">tools</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Food biomass</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M14" display="inline"><mml:mrow><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">kg</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Time allocated to provisioning activity</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M15" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M16" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">d</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Time allocated to other activity</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M17" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M18" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">d</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Experienced affect</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M19" display="inline"><mml:mrow><mml:msub><mml:mi>X</mml:mi><mml:mtext>affect</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">unitless</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup><?xmltex \end{scaleboxenv}?></oasis:table></table-wrap>

      <?xmltex \floatpos{t}?><fig id="Ch1.F3"><?xmltex \currentcnt{3}?><?xmltex \def\figurename{Figure}?><label>Figure 3</label><caption><p id="d1e945">Model architecture, shown within the conceptual framework of Fig. 2. Dependencies are shown as arrows: (i) food supply vs. metabolic demand influences the time allocated to provisioning, <inline-formula><mml:math id="M20" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>; (ii) time allocation to provisioning and the availability of provisioning Things <inline-formula><mml:math id="M21" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> influence the per capita extraction rate of edible food <inline-formula><mml:math id="M22" display="inline"><mml:mrow><mml:msub><mml:mi>E</mml:mi><mml:mi mathvariant="normal">food</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>; (iii) the extraction rate of <inline-formula><mml:math id="M23" display="inline"><mml:mrow><mml:msub><mml:mi>E</mml:mi><mml:mi mathvariant="normal">food</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> determines the change in human biomass <inline-formula><mml:math id="M24" display="inline"><mml:mrow><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>; (iv) time allocation between activities <inline-formula><mml:math id="M25" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M26" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> influences population affect <inline-formula><mml:math id="M27" display="inline"><mml:mrow><mml:msub><mml:mi>X</mml:mi><mml:mtext>affect</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>; (v) time allocation influences the relative formation rates of neural structure types, either <inline-formula><mml:math id="M28" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> or <inline-formula><mml:math id="M29" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>.</p></caption>
          <?xmltex \igopts{width=236.157874pt}?><graphic xlink:href="https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021-f03.png"/>

        </fig>

      <p id="d1e1065"><?xmltex \hack{\newpage}?>The entire planet is represented here as a single entity without any spatial
resolution, for ease of illustration, similar to the seminal model by
<xref ref-type="bibr" rid="bib1.bibx25" id="text.53"/>. This “zero-dimensional” spatial form is common for
a proof-of-concept in Earth system modeling (e.g. <xref ref-type="bibr" rid="bib1.bibx68" id="altparen.54"/>),
though it is envisioned that the most useful application for this framework
would be on a global grid. The model is discrete in time and is solved
numerically through finite differences from a prescribed initial state using
forward time steps of 1 week.</p>
</sec>
<?pagebreak page679?><sec id="Ch1.S5.SS3">
  <label>5.3</label><title>Process overview and scheduling</title>
      <p id="d1e1083">First, the population motivational factors are calculated, based on the food
surplus and time allocation from the prior time step, and these motivational
factors are used to calculate the time allocation for the current
time step. Next, the mass flux of provisioned food is calculated as a function
of the biomass variables (<inline-formula><mml:math id="M30" display="inline"><mml:mrow><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M31" display="inline"><mml:mrow><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>) and the
time allocation to provisioning (<inline-formula><mml:math id="M32" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>), and the biomass
is updated to conserve mass while accounting for metabolism. Finally, the
neural characteristics are updated, and the subjective experience is recorded,
according to the time allocation.</p>
</sec>
<sec id="Ch1.S5.SS4">
  <label>5.4</label><title>Principles</title>
      <p id="d1e1128">The model considers a few physical features of a global human population, tied
together by a novel use of time allocation as the central dynamical
variable. Two activities are defined in terms of their physical outcomes (the
“what” of the activities). The allocation of available time between the two
activities is determined by the relative strengths of their motivational
factors within the population.</p>
      <p id="d1e1131">The first activity, <italic>provision</italic>, includes all activities required to extract
edible organic matter from the environment and distribute it to the population
in an edible form. Conceptually this could include hunting, fishing, or
farming, as well as any necessary processing and transportation of food. The
transfer of biomass between edible food and the human population is strictly
conserved, and the model uses realistic values for human metabolic and growth
rates. The second activity, <italic>other</italic>, includes all other non-essential
activities in which the population is engaged.</p>
      <p id="d1e1140">Motivational factors are conceptualized as originating from the individual
level (at which state-dependent neural activation would occur), scaled up to
population-level changes in time allocation as modulated by social
structures. Motivations are formulated as competing influences on time
allocation, similar to the dynamics-of-action approach used by
<xref ref-type="bibr" rid="bib1.bibx7" id="text.55"/> but applied here at a population level rather than an individual level. I am not aware of prior works that use the same formulation,
but the precise means by which this is achieved are not the focus
here. Rather, the goal is simply to simulate internally consistent dynamical
links.</p>
      <p id="d1e1146">A simple model is used for changes in the neural structure of the population,
based on two simplifying assumptions: first, that the rate of new synapse
formation within the population is randomly distributed at a constant rate
within individual cortexes and, second, that synapses that are being fired
through activation are more likely to strengthen and persist
<xref ref-type="bibr" rid="bib1.bibx6" id="paren.56"/>. Under these two assumptions, the development of
strong synapses, which then become important pathways for future thoughts, are
dependent on engagement in relevant activities. In this way, the time
allocation to activities contributes to the modification of the neural structure.</p>
      <p id="d1e1153"><?xmltex \hack{\newpage}?>Finally, one metric of subjective well-being is used here: the affect balance
associated with different activities. It is assumed that a population-average
level of affect <inline-formula><mml:math id="M33" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> occurs under each activity <inline-formula><mml:math id="M34" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula>, as determined by
many factors that are not resolved here. Basically, one of the activities is
bound to be more enjoyable than the other. Because provisioning generally
falls under the category of work, whether or not it is done through the formal
economy, it is assumed to incur a lower level of affect. The <italic>other</italic> activity,
although sure to include many sub-activities that are unpleasant, is assumed
to incur a higher overall average affect. Note that this analysis ignores any
sense of eudaemonia that may result from either activity and is purely
hedonic.</p>
</sec>
<sec id="Ch1.S5.SS5">
  <label>5.5</label><title>Emergence</title>
      <p id="d1e1186">The model generates emergent outcomes through the integration of the
state-dependent equations. These emergent features include the temporal
evolution of food biomass, human population size, time allocation, neural
structure, and experience. The details of these features are dependent on
parameter choices, which were not exhaustively explored here.</p>
</sec>
<sec id="Ch1.S5.SS6">
  <label>5.6</label><title>Adaptation</title>
      <p id="d1e1197">The human population dynamically allocates the available time between the two
available activities. This allocation is determined by the motivation to
provision food, which is a function of the food supply rate relative to the
population metabolic requirements, in competition with the motivation to do
something other than provisioning food.</p>
      <p id="d1e1200">Human behaviour is exceedingly complex and cannot be predicted from first
principles. Here, the motivational responses at the population scale are
approximated by smooth response functions, reflecting competing tendencies to
alter time allocation between activities in response to changes in relevant
state variables. The simple model considers two motivational factors – <inline-formula><mml:math id="M35" display="inline"><mml:mrow><mml:msub><mml:mi>m</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M36" display="inline"><mml:mrow><mml:msub><mml:mi>m</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> – each of which has a value
between 0 and 1, indicating the relative drive to increase in the fraction of
time devoted to the corresponding activity. To represent saturating responses
to an input variable, the Holling type 2 formulation is used, since it
provides stability and each usage introduces only one additional parameter
(<inline-formula><mml:math id="M37" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula>). Each motivational factor is then weighted by a response coefficient
<inline-formula><mml:math id="M38" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula>, which reflects the strength with which time is reallocated to the
activity in response to the motivational factor.</p>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T2" specific-use="star"><?xmltex \currentcnt{2}?><label>Table 2</label><caption><p id="d1e1242">Model parameters.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="3">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="left"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Variable</oasis:entry>
         <oasis:entry colname="col2">Symbol</oasis:entry>
         <oasis:entry colname="col3">Default value</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">Net primary production</oasis:entry>
         <oasis:entry colname="col2">NPP</oasis:entry>
         <oasis:entry colname="col3">54 <inline-formula><mml:math id="M39" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Pg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">C</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">yr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Edible fraction</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M40" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ϕ</mml:mi><mml:mtext>edible</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M41" display="inline"><mml:mn mathvariant="normal">0.005</mml:mn></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Human maximum growth rate</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M42" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>max</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M43" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.03</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">yr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Human metabolism</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M44" display="inline"><mml:mi mathvariant="italic">ω</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M45" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.02</mml:mn><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Initial mass per human</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M46" display="inline"><mml:mi mathvariant="italic">β</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M47" display="inline"><mml:mrow><mml:mn mathvariant="normal">6</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">C</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">human</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Food distribution parameter</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M48" display="inline"><mml:mi mathvariant="italic">σ</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M49" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.014</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:msub><mml:mi mathvariant="normal">kg</mml:mi><mml:mi mathvariant="normal">food</mml:mi></mml:msub><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msubsup><mml:mi mathvariant="normal">kg</mml:mi><mml:mi mathvariant="normal">human</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msubsup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Available time</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M50" display="inline"><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mtext>avail</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M51" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.6</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:mi mathvariant="normal">d</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Sensitivity to food shortage</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M52" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M53" display="inline"><mml:mn mathvariant="normal">0.4</mml:mn></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Sensitivity to other time shortage</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M54" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M55" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.3</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:mi mathvariant="normal">d</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Reactivity to food shortage</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M56" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M57" display="inline"><mml:mn mathvariant="normal">1</mml:mn></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Reactivity to other shortage</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M58" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M59" display="inline"><mml:mn mathvariant="normal">1</mml:mn></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Maximum efficiency of provisioning</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M60" display="inline"><mml:mrow><mml:msubsup><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mtext>provision</mml:mtext><mml:mtext>max</mml:mtext></mml:msubsup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M61" display="inline"><mml:mrow><mml:mn mathvariant="normal">3</mml:mn><mml:mo>×</mml:mo><mml:msup><mml:mn mathvariant="normal">10</mml:mn><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">14</mml:mn></mml:mrow></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mrow class="unit"><mml:msubsup><mml:mi mathvariant="normal">kg</mml:mi><mml:mi mathvariant="normal">human</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msubsup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Provisioning Things half-saturation constant</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M62" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mrow><mml:mi>T</mml:mi><mml:mtext>provision</mml:mtext></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M63" display="inline"><mml:mrow><mml:mn mathvariant="normal">50</mml:mn><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mrow class="unit"><mml:msub><mml:mi mathvariant="normal">kg</mml:mi><mml:mrow><mml:mi>T</mml:mi><mml:mi mathvariant="normal">provision</mml:mi></mml:mrow></mml:msub><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">human</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Food decay rate</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M64" display="inline"><mml:mi mathvariant="italic">λ</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M65" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.05</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Normalized synapse formation rate</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M66" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>synapse</mml:mtext></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M67" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">75</mml:mn><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">human</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">yr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Synapse destruction rate</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M68" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">λ</mml:mi><mml:mtext>synapse</mml:mtext></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M69" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">yr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mrow></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Affect during provisioning activity</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M70" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M71" display="inline"><mml:mn mathvariant="normal">0.2</mml:mn></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Affect during other activity</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M72" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M73" display="inline"><mml:mn mathvariant="normal">0.5</mml:mn></mml:math></inline-formula></oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d1e1933">The <inline-formula><mml:math id="M74" display="inline"><mml:mi>r</mml:mi></mml:math></inline-formula> and <inline-formula><mml:math id="M75" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula> parameters are intended to reflect the combined outcomes of
individual psychology and societal processes (where societal processes include
all cultural, social, political, and economic interactions). Neither of these
parameters has a direct equivalence that can be measured precisely, a common
occurrence in ecological modeling, and their values are chosen in order to
produce reasonable model behaviour.  Although they are held constant in the
individual<?pagebreak page680?> simulations shown here, they would not in reality be static
properties of the population but could in theory be dynamically linked to
neural structure state variables. However this would go beyond the simple
scope of the current illustration.</p>
</sec>
<sec id="Ch1.S5.SS7">
  <label>5.7</label><title>Initialization</title>
      <p id="d1e1958">Initial values for state variables were chosen to ensure stable
integration. These initial values included a small human population and large
food biomass, in order to allow the population to grow continually for a
couple of centuries under typical parameter values. Parameter values are given
in Table 2.</p>
</sec>
<sec id="Ch1.S5.SS8">
  <label>5.8</label><title>Input data</title>
      <p id="d1e1969">The model does not use input data to represent time-varying processes.</p>
</sec>
<sec id="Ch1.S5.SS9">
  <label>5.9</label><title>Submodels</title>
<sec id="Ch1.S5.SS9.SSS1">
  <label>5.9.1</label><title>Food shortage</title>
      <p id="d1e1987">The mass-specific population average food shortage (<inline-formula><mml:math id="M76" display="inline"><mml:mrow><mml:msub><mml:mtext>shortage</mml:mtext><mml:mtext>ave</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>)
is calculated as the difference between the total provisioned food for the
time step and the food required to meet metabolic needs plus additional
growth:

                  <disp-formula id="Ch1.E1" content-type="numbered"><label>1</label><mml:math id="M77" display="block"><mml:mstyle class="stylechange" displaystyle="true"/><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mtext>shortage</mml:mtext><mml:mtext>ave</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">ω</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>)</mml:mo><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub><mml:mo>-</mml:mo><mml:mtext>provisioned</mml:mtext><mml:mo>)</mml:mo><mml:mo>/</mml:mo><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub><mml:mo>.</mml:mo></mml:mrow></mml:math></disp-formula>

            In order to illustrate how population state variables can capture
within-population variations, the available food is assumed to be unequally
partitioned within the population following a normal distribution about the
average shortage, characterized by a standard deviation <inline-formula><mml:math id="M78" display="inline"><mml:mi mathvariant="italic">σ</mml:mi></mml:math></inline-formula>. The
cumulative distribution function provides the fraction of the population
<inline-formula><mml:math id="M79" display="inline"><mml:mrow><mml:msub><mml:mtext>shortage</mml:mtext><mml:mtext>frac</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> that would experience some level of food shortage, as
illustrated for two values of <inline-formula><mml:math id="M80" display="inline"><mml:mi mathvariant="italic">σ</mml:mi></mml:math></inline-formula> in Fig. 4. The prevalence of
experienced food shortage is assumed to drive the response of time allocation,
according to the motivational parameters.</p>
</sec>
<sec id="Ch1.S5.SS9.SSS2">
  <label>5.9.2</label><title>Motivations</title>
      <p id="d1e2082">The motivation to allocate time to provisioning is a function of the food
shortage, given by

                  <disp-formula id="Ch1.E2" content-type="numbered"><label>2</label><mml:math id="M81" display="block"><mml:mstyle displaystyle="true" class="stylechange"/><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi>m</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mtext>shortage</mml:mtext><mml:mtext>frac</mml:mtext></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mtext>food</mml:mtext></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mtext>shortage</mml:mtext><mml:mtext>frac</mml:mtext></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>.</mml:mo></mml:mrow></mml:math></disp-formula></p>
      <p id="d1e2122">The value of the half-saturation constant <inline-formula><mml:math id="M82" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> determines the
relative strength with which the population is motivated to respond to a given
shortage, with a smaller value responding more strongly to smaller
undernourished fractions and saturating more quickly (Fig. 4).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F4" specific-use="star"><?xmltex \currentcnt{4}?><?xmltex \def\figurename{Figure}?><label>Figure 4</label><caption><p id="d1e2138">Food shortage and provisioning motivation. Panel <bold>(a)</bold> shows the fraction of population experiencing food shortage for two values of <inline-formula><mml:math id="M83" display="inline"><mml:mi mathvariant="italic">σ</mml:mi></mml:math></inline-formula>. A larger value of <inline-formula><mml:math id="M84" display="inline"><mml:mi mathvariant="italic">σ</mml:mi></mml:math></inline-formula> corresponds to greater inequality (red) compared to the value used in the simulations (blue). Panel <bold>(b)</bold> shows the population motivation factor for provisioning as a function of the shortage fraction. The relative motivation strength is shown for highly responsive (low <inline-formula><mml:math id="M85" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula>, purple) and weakly responsive (high <inline-formula><mml:math id="M86" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula>, yellow) societies.</p></caption>
            <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021-f04.png"/>

          </fig>

      <p id="d1e2183">Obtaining food is a primary concern for all animals, but they also tend to
spend some fraction of time doing other things. Depending on the species, they
might invest time developing burrows or nests, engaging in courtship and
mating, or resting in a safe place. Humans, more than any other animal, are
characterized by the wide range of activities in which they are motivated to
engage, other than obtaining food. The associated motivational factor is
termed here <inline-formula><mml:math id="M87" display="inline"><mml:mrow><mml:msub><mml:mi>m</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> and assumed to increase in intensity as the
time allocated to <inline-formula><mml:math id="M88" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> is decreased. This motivation could be
construed at an individual level, such as the individual desire to<?pagebreak page681?> do
something more enjoyable, rewarding, or relaxing than food provision. It could
also occur through social mechanisms, such as cultural norms to engage in
rituals or constructing religious buildings, or through economic mechanisms
such as a decrease in provision labour wages as food availability is
increased. No attempt is made here to represent these mechanisms (i.e. the
how); instead all are simply bundled in a non-provision motivation, given
by the following equation:

                  <disp-formula id="Ch1.E3" content-type="numbered"><label>3</label><mml:math id="M89" display="block"><mml:mstyle displaystyle="true" class="stylechange"/><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi>m</mml:mi><mml:mtext>other</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mo>∑</mml:mo><mml:mi>i</mml:mi></mml:msub><mml:mi>A</mml:mi><mml:mo>-</mml:mo><mml:msub><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mtext>other</mml:mtext></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mo>∑</mml:mo><mml:mi>i</mml:mi></mml:msub><mml:mi>A</mml:mi><mml:mo>-</mml:mo><mml:msub><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>

            where <inline-formula><mml:math id="M90" display="inline"><mml:mrow><mml:msub><mml:mo>∑</mml:mo><mml:mi>i</mml:mi></mml:msub><mml:mi>A</mml:mi></mml:mrow></mml:math></inline-formula> is the total time available for the defined activities and
<inline-formula><mml:math id="M91" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> represents the rate at which <inline-formula><mml:math id="M92" display="inline"><mml:mrow><mml:msub><mml:mi>m</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> approaches
saturation with increasing time spent provisioning.</p>
</sec>
<sec id="Ch1.S5.SS9.SSS3">
  <label>5.9.3</label><title>Time allocation</title>
      <p id="d1e2308">Each of the time allocation terms <inline-formula><mml:math id="M93" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mi>i</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is constrained to vary between 0 and
<inline-formula><mml:math id="M94" display="inline"><mml:mrow><mml:msub><mml:mo>∑</mml:mo><mml:mi>i</mml:mi></mml:msub><mml:mi>A</mml:mi></mml:mrow></mml:math></inline-formula>. For the illustrative model here, unresolved essential activities
(e.g. sleep, meals, personal hygiene) are assumed to occupy 40 <inline-formula><mml:math id="M95" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">%</mml:mi></mml:mrow></mml:math></inline-formula> of
the total time, so that <inline-formula><mml:math id="M96" display="inline"><mml:mrow><mml:msub><mml:mo>∑</mml:mo><mml:mi>i</mml:mi></mml:msub><mml:mi>A</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.6</mml:mn></mml:mrow></mml:math></inline-formula>. The model is constructed to place
multiple motivations in competition for the remaining available time within
the population. As such, the competing motivations exist in tension with each
other, and the outcome represents a dynamic balance between them. The changes
in motivated time allocation are given by

                  <disp-formula specific-use="align" content-type="numbered"><mml:math id="M97" display="block"><mml:mtable displaystyle="true"><mml:mlabeledtr id="Ch1.E4"><mml:mtd><mml:mtext>4</mml:mtext></mml:mtd><mml:mtd><mml:mstyle class="stylechange" displaystyle="true"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:msubsup><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext><mml:mo>′</mml:mo></mml:msubsup></mml:mrow><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:mi>t</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:msub><mml:mi>r</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:msub><mml:mi>m</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="Ch1.E5"><mml:mtd><mml:mtext>5</mml:mtext></mml:mtd><mml:mtd><mml:mstyle class="stylechange" displaystyle="true"/></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:msubsup><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext><mml:mo>′</mml:mo></mml:msubsup></mml:mrow><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:mi>t</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub><mml:msub><mml:mi>m</mml:mi><mml:mtext>other</mml:mtext></mml:msub><mml:mo>.</mml:mo></mml:mrow></mml:mtd></mml:mlabeledtr></mml:mtable></mml:math></disp-formula></p>
      <p id="d1e2443">Thus, the motivational factors act upon the previously allocated time. The
individual motivated time allocation terms <inline-formula><mml:math id="M98" display="inline"><mml:mrow><mml:msubsup><mml:mi>A</mml:mi><mml:mi>i</mml:mi><mml:mo>′</mml:mo></mml:msubsup></mml:mrow></mml:math></inline-formula> are then divided by the sum
of all <inline-formula><mml:math id="M99" display="inline"><mml:mrow><mml:msubsup><mml:mi>A</mml:mi><mml:mi>i</mml:mi><mml:mo>′</mml:mo></mml:msubsup></mml:mrow></mml:math></inline-formula> and multiplied by <inline-formula><mml:math id="M100" display="inline"><mml:mrow><mml:msub><mml:mo>∑</mml:mo><mml:mi>i</mml:mi></mml:msub><mml:mi>A</mml:mi></mml:mrow></mml:math></inline-formula>, distributing the available time
among the possible activities according to their strength of motivation, while
respecting the conservation of total time.</p>
</sec>
<sec id="Ch1.S5.SS9.SSS4">
  <label>5.9.4</label><title>Effectiveness multiplier</title>
      <p id="d1e2493">The provisioning activity is associated with a multiplier
<inline-formula><mml:math id="M101" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> that represents how effectively the activity is
carried out by the population. This effectiveness multiplier could reflect the
culturally transmitted knowledge and skill within a population, as well as the
availability of tools and machinery, the quality of those tools and machinery,
and non-human aspects of the activity.  The multiplier has units of
<inline-formula><mml:math id="M102" display="inline"><mml:mrow class="unit"><mml:mo>(</mml:mo><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msub><mml:mi mathvariant="normal">C</mml:mi><mml:mtext>human</mml:mtext></mml:msub><mml:msup><mml:mo>)</mml:mo><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>, representing the fraction of available
edible material that would be provisioned per human time. The value of
<inline-formula><mml:math id="M103" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> increases with the per capita mass of
provisioning Things, <inline-formula><mml:math id="M104" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>, approaching a maximum that
reflects the saturation of the active population with available tools. This
saturation is analogous to the saturation of enzymes with reagents, so the
Michaelis–Menten or Holling type 2 formulation is also used here, so that

                  <disp-formula id="Ch1.E6" content-type="numbered"><label>6</label><mml:math id="M105" display="block"><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><?xmltex \hack{\hbox\bgroup\fontsize{8.9}{8.9}\selectfont$\displaystyle}?><mml:msub><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:msubsup><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mtext>provision</mml:mtext><mml:mtext>max</mml:mtext></mml:msubsup><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:mo>/</mml:mo><mml:mo>(</mml:mo><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub><mml:mo>/</mml:mo><mml:mi mathvariant="italic">β</mml:mi><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mrow><mml:mi>T</mml:mi><mml:mtext>provision</mml:mtext></mml:mrow></mml:msub><mml:mo>+</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:mo>/</mml:mo><mml:mo>(</mml:mo><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub><mml:mo>/</mml:mo><mml:mi mathvariant="italic">β</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>.</mml:mo><?xmltex \hack{$\egroup}?></mml:mrow></mml:math></disp-formula></p>
      <p id="d1e2650">In these simulations, <inline-formula><mml:math id="M106" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> is held at a constant per capita
value.</p>
</sec>
<sec id="Ch1.S5.SS9.SSS5">
  <label>5.9.5</label><title>Mass fluxes</title>
      <p id="d1e2673">The dynamical change in edible biomass <inline-formula><mml:math id="M107" display="inline"><mml:mrow><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> is given by

                  <disp-formula id="Ch1.E7" content-type="numbered"><label>7</label><mml:math id="M108" display="block"><mml:mrow><?xmltex \hack{\hbox\bgroup\fontsize{8.9}{8.9}\selectfont$\displaystyle}?><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:mi>t</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:msub><mml:mi mathvariant="italic">ϕ</mml:mi><mml:mtext>edible</mml:mtext></mml:msub><mml:mtext>NPP</mml:mtext><mml:mo>-</mml:mo><mml:mi mathvariant="italic">λ</mml:mi><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub><mml:msub><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub><mml:mo>.</mml:mo><?xmltex \hack{$\egroup}?></mml:mrow></mml:math></disp-formula></p>
      <?pagebreak page682?><p id="d1e2752">The fraction of net primary productivity (NPP) that is allocated to edible material,
<inline-formula><mml:math id="M109" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ϕ</mml:mi><mml:mtext>edible</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>, would vary with many factors including the ecosystem
type, climate, and human agency. For example, human activity can modify
<inline-formula><mml:math id="M110" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ϕ</mml:mi><mml:mtext>edible</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>, increasing it through deliberate modifications
including agriculture and aquaculture or decreasing it by destructively
harvesting and overhunting or overfishing. Human activity could also modify NPP,
increasing it by changing vegetation to more productive varieties or by
fertilizing and irrigating or decreasing it by causing soil erosion, nutrient
loss, or other forms of ecological degradation. For simplicity, all of these
factors are conceptually bundled within a constant value of
<inline-formula><mml:math id="M111" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ϕ</mml:mi><mml:mtext>edible</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> as modifications of edible NPP relative to a
“pristine” state (i.e. untouched by humans). For reference, the present-day
global annual production of edible material is approximately 0.9 <inline-formula><mml:math id="M112" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Pg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>
(the mass of carbon within all edible primary crops, processed crops, and
animal products) according to the analysis of <xref ref-type="bibr" rid="bib1.bibx2" id="text.57"/>,
implying a global <inline-formula><mml:math id="M113" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ϕ</mml:mi><mml:mtext>edible</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> of roughly 0.02 for a global NPP of
54 <inline-formula><mml:math id="M114" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Pg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx50" id="paren.58"/>.</p>
      <p id="d1e2828">In the second term, <inline-formula><mml:math id="M115" display="inline"><mml:mi mathvariant="italic">λ</mml:mi></mml:math></inline-formula> (<inline-formula><mml:math id="M116" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>) represents the consumption of
potentially edible material by all non-human organisms such as other mammals,
birds, insects, fungi, or bacteria. This non-human consumption is assumed to be
first order with respect to <inline-formula><mml:math id="M117" display="inline"><mml:mrow><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>, for simplicity. The decay
constant would be expected to vary with food type and environment but would
generally be on the order of weeks.</p>
      <p id="d1e2863">The final term represents the collection and essential processing of edible
material by humans, which is the outcome of the provisioning activity. The
term depends on <inline-formula><mml:math id="M118" display="inline"><mml:mrow><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>, the size of the human population,
<inline-formula><mml:math id="M119" display="inline"><mml:mrow><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M120" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msub><mml:mi mathvariant="normal">C</mml:mi><mml:mi mathvariant="normal">human</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), the fraction of time allocated to
provisioning, <inline-formula><mml:math id="M121" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M122" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">d</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>), and the effectiveness with
which the population provisions the existing edible material per unit time
<inline-formula><mml:math id="M123" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M124" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msubsup><mml:mi mathvariant="normal">C</mml:mi><mml:mi mathvariant="normal">human</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msubsup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>). The use of
linear dependences is sure to be inappropriate, given that optimal resources
will be harvested first, and diminishing returns would be expected to lead to
a sublinear dependence on <inline-formula><mml:math id="M125" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (note this
is equivalent to labour in the similar Cobb–Douglas production function,
which typically has an exponent <inline-formula><mml:math id="M126" display="inline"><mml:mrow><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>; <xref ref-type="bibr" rid="bib1.bibx16" id="altparen.59"/>). This
approach could potentially be improved upon in future.</p>
      <p id="d1e3000">Next, the human capacity for food ingestion is given by the product of the
human population <inline-formula><mml:math id="M127" display="inline"><mml:mrow><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> and the sum of the population average
biomass-specific metabolic rates <inline-formula><mml:math id="M128" display="inline"><mml:mi mathvariant="italic">ω</mml:mi></mml:math></inline-formula> and the potential net growth rate
<inline-formula><mml:math id="M129" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula>. Any excess of food provisioned beyond this limit is assumed
to be discarded. An average value of <inline-formula><mml:math id="M130" display="inline"><mml:mi mathvariant="italic">ω</mml:mi></mml:math></inline-formula> is calculated assuming a per capita
energetic requirement of 10 <inline-formula><mml:math id="M131" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">MJ</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> and food energy content of 30 <inline-formula><mml:math id="M132" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kJ</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">g</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">C</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx2" id="paren.60"/>. The value of the maximum growth rate
<inline-formula><mml:math id="M133" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>max</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> – the population growth rate when the rate of food
provisioning is non-limiting – influences the transient behaviour of the model
but not the steady-state outcome. Because <inline-formula><mml:math id="M134" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>max</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> is the maximum
net growth rate, equal to the birth rate minus the death rate (for constant
individual body size), its value reflects both the fertility rates of the
population and the mortality due to disease, violent deaths, and old age. The
fertility rate is dependent on cultural and societal characteristics, while
the rate of death depends on cultural and societal characteristics as well as
exposure to pathogens. Because the cultural and societal aspects of both
fertility and mortality are complex, the model simply considers how the net
result decreases below the potential maximum when assuming zero growth among
the fraction of the population experiencing a food shortage, so that <inline-formula><mml:math id="M135" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msub><mml:mtext>shortage</mml:mtext><mml:mtext>frac</mml:mtext></mml:msub><mml:mo>)</mml:mo><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>max</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>. Food waste is not treated
explicitly but could be regarded as an implicit component of <inline-formula><mml:math id="M136" display="inline"><mml:mi mathvariant="italic">ω</mml:mi></mml:math></inline-formula>,
along with egested food.</p>
      <p id="d1e3133">The human biomass then varies as

                  <disp-formula id="Ch1.E8" content-type="numbered"><label>8</label><mml:math id="M137" display="block"><mml:mrow><?xmltex \hack{\hbox\bgroup\fontsize{8.9}{8.9}\selectfont$\displaystyle}?><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:mi>t</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:mo movablelimits="false">min⁡</mml:mo><mml:mo>(</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>,</mml:mo><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub><mml:msub><mml:mi mathvariant="italic">ϵ</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:msub><mml:mi>E</mml:mi><mml:mtext>food</mml:mtext></mml:msub><mml:mo>-</mml:mo><mml:mi mathvariant="italic">ω</mml:mi><mml:mo>)</mml:mo><mml:mo>)</mml:mo><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub><mml:mo>.</mml:mo><?xmltex \hack{$\egroup}?></mml:mrow></mml:math></disp-formula></p>
</sec>
<sec id="Ch1.S5.SS9.SSS6">
  <label>5.9.6</label><title>Neural structure</title>
      <p id="d1e3212">The basic dynamic by which time allocation modifies the neural structure is
crudely approximated here by

                  <disp-formula id="Ch1.E9" content-type="numbered"><label>9</label><mml:math id="M138" display="block"><mml:mstyle class="stylechange" displaystyle="true"/><mml:mrow><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:msub><mml:mi>N</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:mi mathvariant="normal">d</mml:mi><mml:mi>t</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:msubsup><mml:mi mathvariant="italic">μ</mml:mi><mml:mi>x</mml:mi><mml:mtext>synapse</mml:mtext></mml:msubsup><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub><mml:mo>-</mml:mo><mml:msup><mml:mi mathvariant="italic">λ</mml:mi><mml:mtext>synapse</mml:mtext></mml:msup><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msub><mml:mi>A</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:msub><mml:mi>N</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>

            where <inline-formula><mml:math id="M139" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the number of synapses in the population associated with
activity <inline-formula><mml:math id="M140" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> (normalized to the lifetime synapse production of an average
individual), <inline-formula><mml:math id="M141" display="inline"><mml:mrow><mml:msubsup><mml:mi mathvariant="italic">μ</mml:mi><mml:mi>x</mml:mi><mml:mtext>synapse</mml:mtext></mml:msubsup></mml:mrow></mml:math></inline-formula> is the biomass-specific growth rate of
new synapses that can be activated by activity <inline-formula><mml:math id="M142" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula>, and <inline-formula><mml:math id="M143" display="inline"><mml:mi mathvariant="italic">λ</mml:mi></mml:math></inline-formula> is the
synapse-specific rate of synapse destruction (arbitrarily chosen to provide a
1-year <inline-formula><mml:math id="M144" display="inline"><mml:mi>e</mml:mi></mml:math></inline-formula>-folding timescale, for illustration).</p>
      <p id="d1e3332">In this formulation, synapses are defined by their associated activity. This
does not imply that they are exclusively related to the functional core of the
activity but simply that they are activated and thereby strengthened during
the activity. There could also be overlap between the neural structures of
different activities due to commonalities, which are not resolved here.</p>
      <p id="d1e3335">It is essential that this quantification says nothing about the functional
utility of the structural changes. Many of the accumulated synapses may
contribute little or even be deleterious. The processes by which the brain
selects and amplifies the functional utility of certain synaptic
modifications, while dampening others, remains an important topic of research
in neuroscience <xref ref-type="bibr" rid="bib1.bibx47" id="paren.61"/>. Nonetheless, the fact that
synapses are strengthened in response to activation is well-established
<xref ref-type="bibr" rid="bib1.bibx6" id="paren.62"/>, and it is expected that future work can improve on
this crude representation.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F5" specific-use="star"><?xmltex \currentcnt{5}?><?xmltex \def\figurename{Figure}?><label>Figure 5</label><caption><p id="d1e3347">Time series of a typical model experiment. In <bold>(f)</bold>, the dashed line indicates the metabolic cost of maintaining the population (i.e. <inline-formula><mml:math id="M145" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ω</mml:mi><mml:mtext>human</mml:mtext></mml:msub><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>) and the shaded green area represents the food surplus.</p></caption>
            <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021-f05.png"/>

          </fig>

</sec>
<sec id="Ch1.S5.SS9.SSS7">
  <label>5.9.7</label><title>Subjective state</title>
      <?pagebreak page683?><p id="d1e3383">The instantaneous average affect of the population at time <inline-formula><mml:math id="M146" display="inline"><mml:mi>t</mml:mi></mml:math></inline-formula> is given by the time-weighted mean of the activity-specific affects:

                  <disp-formula id="Ch1.E10" content-type="numbered"><label>10</label><mml:math id="M147" display="block"><mml:mstyle class="stylechange" displaystyle="true"/><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi>x</mml:mi><mml:mtext>affect</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mtext>avail</mml:mtext></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo movablelimits="false">∑</mml:mo><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:msub><mml:mi>A</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>

            which can be rewritten for this two-activity model as
<?xmltex \hack{\newpage}?><?xmltex \hack{\vspace*{-6mm}}?>

                  <disp-formula id="Ch1.E11" content-type="numbered"><label>11</label><mml:math id="M148" display="block"><mml:mstyle class="stylechange" displaystyle="true"/><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:msub><mml:mi>x</mml:mi><mml:mtext>affect</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mtext>other</mml:mtext></mml:msub><mml:mo>-</mml:mo><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mtext>other</mml:mtext></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mtext>provision</mml:mtext></mml:msub><mml:mo>)</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>f</mml:mi><mml:mtext>avail</mml:mtext></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>

            giving a linear decrease with <inline-formula><mml:math id="M149" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> below a maximum affect <inline-formula><mml:math id="M150" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>.</p>
</sec>
</sec>
</sec>
<sec id="Ch1.S6">
  <label>6</label><title>Model analysis and discussion</title>
      <p id="d1e3523">The transient dynamics of the model can assume many forms depending on the
input parameters as a result of the nonlinear interactions between the
societal aspects (motivational factors) and the human–ecosystem coupling (food
provisioning). The following sections describe typical features of the
transient behaviour that are robust across a breadth of reasonable parameter
choices.</p>
<sec id="Ch1.S6.SS1">
  <label>6.1</label><title>Approach to steady-state population</title>
      <p id="d1e3533">When initialized from a population density well below the steady-state value,
the human population grows near-exponentially (Fig. 5a). The food biomass is
drawn down (Fig. 5b), generating decreasing yields for the same effort
(Fig. 5c). Hunger increases in response (Fig. 5d), which drives a greater
<inline-formula><mml:math id="M151" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (Fig. 5e). The surplus (difference between the solid
and dashed line in Fig. 5f) gradually shrinks, until after a couple of
centuries the surplus reaches the point at which it constrains the growth
rate. At this point the population growth rapidly declines to zero and
<inline-formula><mml:math id="M152" display="inline"><mml:mrow><mml:msub><mml:mi>S</mml:mi><mml:mtext>mass</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> reaches a plateau (Fig. 5a). The transition from growth to
plateau happens more sharply than under logistic growth because the modeled
growth rate remains large even as the food surplus shrinks, and the constraint
of food limitation on growth is imposed abruptly. This could be unrealistic
for populations that have sufficient foresight to slow their growth<?pagebreak page684?> rate in
advance of food limitation but is perhaps realistic for populations in which
reproductive rates do not decline in response to declining food
surpluses. Note that mass is not strictly equivalent to the number of humans,
since the mass per human could change.</p>
</sec>
<sec id="Ch1.S6.SS2">
  <label>6.2</label><?xmltex \opttitle{Dependence of population size on $r_{{\text{provision}}}$ or $r_{{\text{other}}}$}?><title>Dependence of population size on <inline-formula><mml:math id="M153" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> or <inline-formula><mml:math id="M154" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></title>
      <p id="d1e3588">Figure 6 shows the same experiment shown in Fig. 5, as well as a second
experiment in which a single parameter value was changed: <inline-formula><mml:math id="M155" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>
was increased by a factor of 4. This increase reflects a greater motivation
within the population to engage in <inline-formula><mml:math id="M156" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> rather than
<inline-formula><mml:math id="M157" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>. Such a motivation could reflect a desire for leisure,
a societal focus on monumental architecture, or a culture of learning – these
distinctions are not resolved here.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F6" specific-use="star"><?xmltex \currentcnt{6}?><?xmltex \def\figurename{Figure}?><label>Figure 6</label><caption><p id="d1e3626">Time series of two model experiments with different values of <inline-formula><mml:math id="M158" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>.</p></caption>
          <?xmltex \igopts{width=497.923228pt}?><graphic xlink:href="https://esd.copernicus.org/articles/12/671/2021/esd-12-671-2021-f06.png"/>

        </fig>

      <p id="d1e3646">The higher <inline-formula><mml:math id="M159" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (relative to <inline-formula><mml:math id="M160" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>) results in
a smaller population size at steady state (Fig. 6a). This occurs even though
the food shortage experienced by the population is the same at steady state:
the population simply decides to allocate less time to provisioning because
their priority is to engage in other activities. Because they provision less
intensively, the food biomass remains more abundant (Fig. 6b), resulting in a
greater provisioning efficiency (Fig. 6g). The greater allocation of time to
other activities results in a large contrast in the neural structure, with
<inline-formula><mml:math id="M161" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> much greater than <inline-formula><mml:math id="M162" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> in the population
with high <inline-formula><mml:math id="M163" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (Fig. 6i). Additionally, the steady-state affect
is greater with high <inline-formula><mml:math id="M164" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> (Fig. 6j), given the assumption that
other activities provide a higher affect than provisioning. Thus, the high
<inline-formula><mml:math id="M165" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> experiment produces a smaller population of happier people
with a more diverse neural structure.</p><?xmltex \hack{\newpage}?>
</sec>
<sec id="Ch1.S6.SS3">
  <label>6.3</label><title>Golden ages</title>
      <p id="d1e3736">Figure 6 illustrates an interesting nonlinear dynamic, particularly pronounced
in the food-focused population (low <inline-formula><mml:math id="M166" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>). During the initial
population growth phase, <inline-formula><mml:math id="M167" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> remains relatively high, since
food is abundant and hunger is low. This allows the development of
<inline-formula><mml:math id="M168" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>, indicating a more diverse neural structure within the
population, and supports a high level of affect. However, as food limitation
approaches, food shortages increase, and the low <inline-formula><mml:math id="M169" display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> causes the
population activity to shift rapidly to <inline-formula><mml:math id="M170" display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mtext>provision</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>. The
<inline-formula><mml:math id="M171" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mtext>other</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula> is no longer maintained at the high level, and the affect
drops. One could conceive that such a century-timescale transient would be
recalled by a society as having passed through a golden age, such as that
mythologized in ancient Greece <xref ref-type="bibr" rid="bib1.bibx8" id="paren.63"/> and frequently
echoed throughout history.</p>
      <p id="d1e3809">This dynamic does not occur under all parameter combinations, and it should be
borne in mind that the model is very simple. However, it serves to illustrate
a straightforward interaction that would be expected to produce temporary
golden ages with abundant food and time for <italic>other</italic> activities such as
learning, producing art, and building public works.</p>
</sec>
</sec>
<sec id="Ch1.S7" sec-type="conclusions">
  <label>7</label><title>Conclusions</title>
      <p id="d1e3824">The global human system can appear overwhelmingly complex, which has
contributed to the general hesitance to include it within Earth system science on a common footing with the atmosphere, ocean, terrestrial ecosystem, marine ecosystem, and cryosphere. This paper has suggested that the global human system can be more effectively integrated with the remainder of the Earth system through an unremitting focus on physical foundations, including an explicit consideration of the aspect we care most about – the human experience of life.</p>
      <p id="d1e3827">The first part of the paper outlined a set of principles, proposed to form the basis of Earth system economics. It was suggested that, despite the inherent challenges, there is promise in striving for improved physical understanding of essential human features. A physically based understanding is less prone to ambiguity and could circumvent disciplinary barriers, opening new opportunities for dialogue across many fields of human study. Section 4 proposed a framework of state variables to capture the entire human system in a way that is both inclusive and functionally consistent. Time allocation provides a central, quantitative anchor for the otherwise bewildering possible range of human activities, while the Soma, neural structures and Things are recognized as persistent, defining features of the human system. The small number of high-level variable classes is intended to facilitate a synoptic global view, while subdivisions of these classes within the overarching framework can allow resolution of important details. Sections 5 and 6 provided an illustration<?pagebreak page685?> of how a numerical model might be constructed within the framework.</p>
      <p id="d1e3830">The ESE approach could be greatly advanced through further progress in three
key domains in the short term. The first is a better understanding of global human time allocation, including improved theoretical foundations and harmonized multinational datasets. The second is a corresponding mapping of human-created Things that is structurally consistent with the resolved human activities and their biophysical outcomes.  And the third includes insights on the process-oriented relationships that link activity and context to multiple dimensions of subjective experience. In the longer term, an improved set of metrics for neural structures that goes beyond the rudimentary approach used here could open the door to realistically quantifying rates of change for key societal characteristics. In addition, there already exists a wealth of complementary approaches to assessing aspects of the biophysical reality of humans. These range from global spatial hunter–gatherer models <xref ref-type="bibr" rid="bib1.bibx59" id="paren.64"/> to human mobility studies <xref ref-type="bibr" rid="bib1.bibx43" id="paren.65"/>, models of physical labour capacity <xref ref-type="bibr" rid="bib1.bibx22" id="paren.66"/>, and the study of societal metabolism <xref ref-type="bibr" rid="bib1.bibx29 bib1.bibx32" id="paren.67"/>. Combining these and other physically based approaches could yield powerful new insights.</p>
      <p id="d1e3845">The ESE approach is intended to help with thinking across scales and disciplines. Although the latter half of this paper has focused on the use of ESE for numerical modeling, it may prove more useful as a conceptual basis for analyzing the global human system in general. By providing an overarching organizing framework, it is hoped that ESE may help to unite disparate learnings from the social sciences and to bring them together with natural sciences to answer urgent questions about the functioning of the human–Earth system.</p>
</sec>

      
      </body>
    <back><notes notes-type="codeavailability"><title>Code availability</title>

      <p id="d1e3852">All MATLAB code used to run the model and generate figures is available for download from the Zenodo archive at <ext-link xlink:href="https://doi.org/10.5281/zenodo.4660554" ext-link-type="DOI">10.5281/zenodo.4660554</ext-link> <xref ref-type="bibr" rid="bib1.bibx26" id="paren.68"/>.</p>
  </notes><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d1e3864">The contact author has declared that there are no competing interests.</p>
  </notes><ack><title>Acknowledgements</title><p id="d1e3870">I am very grateful to Maria Pastor, Viki Reyes-Garcia, Dan Zhu, Priscilla LeMezo, William Fajzel, Ian Hatton, Sara Miñarro, and all members of the iESD laboratory for insightful and inspiring discussions. Kim Scherrer, Chris Barrington-Leigh, and Jeroen Van Den Bergh provided valuable feedback on the paper. This project has received funding from the European Research Council (ERC) under the European Union's Horizon 2020 research and innovation programme (grant agreement no. 682602).</p></ack><notes notes-type="financialsupport"><title>Financial support</title>

      <p id="d1e3875">This research has been supported by the H2020 European Research Council (grant no. BIGSEA 682602).</p>
  </notes><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d1e3881">This paper was edited by Christian Franzke and reviewed by Marcin Czupryna and one anonymous referee.</p>
  </notes><ref-list>
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<abstract-html><p>The study of humans has largely been carried out in isolation from the study of the non-human Earth system. This isolation has encouraged the development of incompatible philosophical, aspirational, and methodological approaches that have proven very difficult to integrate with those used for the non-human remainder of the Earth system. Here, an approach is laid out for the scientific study of the global human system that is intended to facilitate seamless integration with non-human processes by striving for a consistent physical basis, for which the name Earth system economics is proposed. The approach is typified by a foundation on state variables, central among which is the allocation of time amongst activities by human populations, and an orientation towards considering human experience. A framework is elaborated which parses the Earth system into six classes of state variables, including a neural structure class that underpins many essential features of humanity.  A working example of the framework is then illustrated with a simple numerical model, considering a global population that is engaged in one of two waking activities: provisioning food or doing something else. The two activities are differentiated by their motivational factors, outcomes on state variables, and associated subjective experience. While the illustrative model is a gross simplification of reality, the results suggest how neural characteristics and subjective experience can emerge from model dynamics.  The approach is intended to provide a flexible and widely applicable strategy for understanding the human–Earth system, appropriate for physically based assessments of the past and present, as well as contributing to long-term model projections that are naturally oriented towards improving human well-being.</p></abstract-html>
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